Ecology, management and ecosystem services of forests and agroforestry systems, with particular emphasis on tropical and upland environments. Biodiversity conservation in protection and production forests; invasive woody species. Forest restoration and remediation of degraded and post-industrial land. Sustainable forest management and policy including: climate change mitigation; Reduced Emissions from Deforestation and forest Degradation; reduced impact logging; silviculture; secondary and buffer-zone forest management; community participation; monitoring. Forest ecology including: natural regeneration; resistance and resilience of forests to disturbance; succession; landscape; climate change adaptation.  Ecosystem ecology across a range of forest, agroforestry, agricultural and upland land uses: nutrient and carbon cycling; sustainable intensification.



Rowlands, R.S. Understanding the wider environmental impacts of bracken control methods, including the use of livestock. Co-supervised with Dave Chadwick and Dave Styles

Barton, O. Modelling an expanding roe deer population in Wales (MSc (Res)). Co-supervised with Graeme Shannon

Mosbery, M. Temperate lianes: Forest ecology and urban habitat enrichment.

Govigli, V.M. Modelling socio-ecological interactions of sacred forests in Northern Greece. Co-supervised with John Halley and Jenny Wong



(Papers directly related to climate change mitigation and adaptation are indicated by C)


Tropical forest management, degradation and conservation

Kleinschroth, F., Healey, J.R. Gourlet-Fleury, S., Mortier, F. & Stoica, R.S. (2017). Effects of logging on roadless space in intact forest landscapes of the Congo Basin. Conservation Biology 31, 469-480 (

Kleinschroth, F. & Healey, J.R. (2017). Impacts of logging roads on tropical forests. Biotropica (

Kleinschroth, F., Healey, J.R., Sist, P., Mortier, F. & Gourlet-Fleury, S. (2016). How persistent are the impacts of logging roads on Central African forest vegetation? Journal of Applied Ecology 53, 1127-1137.  DOI: 10.1111/1365-2664.12661. C

Kleinschroth, F., Healey, J.R. & Gourlet-Fleury, S. (2016). Sparing forests in Central Africa: re-use old logging roads to avoid creating new ones. Frontiers in Ecology and the Environment 14, 9-10 (

Kleinschroth, F., Gourlet-Fleury, S., Gond, V., Sist, P. & Healey, J.R. (2016). Logging roads in tropical forests: Synthesis of literature written in French and English highlights environmental impact reduction through improved engineering.  Bois et Forêts des Tropiques 328 (2), 13-26.

Kleinschroth, F., Gourlet-Fleury, S., Sist, P., Mortier, F. & Healey, J.R. (2015). Legacy of logging roads in the Congo Basin: how persistent are the scars in forest cover? Ecosphere 6(4): 64 (

Aerts, R., Van Overtveld, K., Wassie, A., Abiyu, A., Demissew, S., Daye, D.D., Giday, K., Haile, M., TewoldeBerhan, S., Teketay, D., Teklehaimanot, Z., Binggeli, P., Deckers, S., Friis, I., Gratzer, G., Hermy, M., Sterck, F.J., Muys, B., Bongers, F. & Healey, J.R. (2016). Conservation of the Ethiopian church forests: threats, opportunities and implications for their management. Science of the Total Environment 551-552, 404-414.  DOI: 10.1016/j.scitotenv.2016.02.034

Daye, D.D. & Healey, J.R. (2015).  Impacts of land-use change on sacred forests at the landscape scale. Global Ecology and Conservation 3, 349-358 (

Desissa, D. & Helaey, J.R. (2012). Sacred groves in the Gamo Highlands of southern Ethiopia: biocultural values and threats. Langscape 2 (11), 44-47.

Morales-Barquero, L., Borrego, A., Skutsch, M., Kleinn, C. & Healey, J.R. (2015). Identification and quantification of drivers of forest degradation in tropical dry forests: a case study in Western Mexico. Land Use Policy 49, 296-309 ( C

Morales-Barquero, L., Skutsch, M., Jardel-Peláez, E.J., Ghilardi, A., Kleinn, C. & Healey, J.R. (2014). Operationalizing the definition of forest degradation for REDD+, with application to Mexico. Forests 5, 1653-1681 (doi: 10.3390/f507 1653; C

Petrokofsky, G., Sist, P., Blanc, L., Doucet, J.-L., Finegan, B., Gourlet-Fleury, S., Healey, J.R., Livoreil, B., Nasi, R., Peña-Claros, M., Putz, F.E. & Zhou, W. (2015). Comparative effectiveness of silvicultural interventions for increasing timber production and sustaining conservation values in natural tropical production forests. A systematic review protocol. Environmental Evidence 4: 8.  DOI: 10.1186/s13750-015-0034-7. C

Rendón Thompson, O.R., Paavola, J., Healey, J.R., Jones, J.P.G, Baker, T.R. & Torres, J. (2013). Reducing emissions from deforestation and forest degradation (REDD+): transaction costs of six Peruvian projects. Ecology and Society 18(1), 17 ( C

Pullin, A.S., Bangpan, M., Dalrymple, S., Dickson, K., Haddaway, N.R., Healey, J.R., Hauari, H., Hockley, N., Jones, J.P.G., Knight, T., Vigurs, C. & Oliver, S. (2013). Human well-being impacts of terrestrial protected areas. Environmental Evidence 2, 19.

Bowler, D.E., Buyung-Ali, L.M., Healey, J.R., Jones, J.P.G., Knight T.M. & Pullin, A.S. (2012). Does community forest management provide global environmental benefit and improve local welfare? Frontiers in Ecology and the Environment 10, 29-36. C


Tropical forest ecology and restoration

Poorter, L…, Healey, J.R.….. (2015). Diversity enhances carbon storage in tropical forests. Global Ecology and Biogeography 24, 1314-1328.  DOI:10.1111/geb.12364. C

Derroire, G., Balvanera, P., Castellanos-Castro, C., Decocq, G., Kennard, D., Lebrija-Trejos, E., Leiva, J., Oden, P.C., Powers, J., Rico-Gray, V., Tigabu, M. & Healey, J.R. (2016). Resilience of tropical dry forests – a meta-analysis of changes in species diversity and composition during secondary succession. Oikos 125, 1386-1397.  DOI: 10.1111/oik.03229.

Derroire, G., Coe, R. & Healey, J.R. (2016). Isolated trees as nuclei of regeneration in tropical pastures: testing the importance of niche-based and landscape factors. Journal of Vegetation Science 27, 679-691.  DOI: 10.1111/jvs.12404.

Derroire, G., Tigabu, M., Odén, P.C. & Healey, J.R. (2016). The effects of established trees on woody regeneration during secondary succession in tropical dry forests. Biotropica 48, 290-300.  DOI:10.1111/btp.12287.

Tanner, E.V.J., Rodriguez-Sanchez, F., Healey, J.R., Holdaway, R.J. & Bellingham, P.J. (2014). Long-term hurricane damage effects on tropical forest tree growth and mortality. Ecology 95, 2974-2983 (

Chai, S.-L., Healey, J.R. & Tanner, E.V.J. (2012). Evaluation of forest recovery over time and space using permanent plots monitored over 30 years in a Jamaican montane rain forest. PLOS ONE 7(11): e48859.  DOI:10.1371/journal.pone.0048859.



Rahman, S.A., Sunderland, T., Roshetko, J.M. & Healey, J.R. (2017). Facilitating smallholder tree farming in fragmented tropical landscapes: Challenges and potentials for sustainable land management. Journal of Environmental Management 198, 110-121 (

Rahman, S.A., Bredahl Jacobsen, J., Healey, J.R., Roshetko, J.M. & Sunderland, T. (2017). Finding alternatives to swidden agriculture: Does agroforestry improve livelihood options and reduce pressure on existing forest? Agroforestry Systems 91, 185-199.  (

Rahman, S., Sunderland, T., Kshatriya, M., Roshetko, J.M., Pagella, T. & Healey, J.R. (2016). Towards productive landscapes: Trade-offs in tree-cover and income across a matrix of smallholder agricultural land-use systems. Land Use Policy 58, 152-164 (

Rahman, S.A., Sunderland, T., Roshetko, J.M., Basuki, I. & Healey, J.R. (2016). Tree culture of smallholder farmers practicing agroforestry in Gunung Salak Valley, West Java, Indonesia. Small-scale Forestry 58, 152-164 (

Noponen, M.R.A., Healey, J.R., Soto, G. & Haggar, J.P. (2013). Sink or source – the potential of coffee agroforestry systems to sequester atmospheric CO2 into soil organic carbon. Agriculture, Ecosystems & Environment, 175, 60-68. C

Noponen, M.R.A., Haggar, J.P., Edwards-Jones, G., Healey, J.R. (2013). Intensification of coffee systems can increase the effectiveness of REDD mechanisms. Agricultural Systems 119, 1-9.  C

Noponen, M.R.A., Edwards-Jones, G., Haggar, J.P., Soto, G., Attarzadeh, N. & Healey, J.R. (2012). Greenhouse gas emissions in coffee grown with differing input levels under conventional and organic management. Agriculture, Ecosystems & Environment, 151, 6-15.  C


Temperate forest, heathland and peatland ecology

Gray, J. & Healey, J.R. (2016). Overwintering sites of ladybirds, shieldbugs and allied species in Hertfordshire woodland: how important is aspect? Transactions of the Hertfordshire Natural History Society 48 (1), 89-98.

Haddaway, N.R., Burden, A., Evans, C.D., Healey, J.R., Jones, D.L., Dalrymple, S.E. & Pullin, A.S. (2014). Evaluating effects of land management on greenhouse gas fluxes and carbon balances in boreo-temperate lowland peatland systems. Environmental Evidence 2014, 3: 5. DOI: 10.1186/2047-2382-3-5; C

Smith, A.R., Lukac, M., Hood, R., Healey, J.R., Miglietta, F. & Godbold, D.L. (2013). Elevated CO2 enrichment induces a differential biomass response in a mixed species temperate forest plantation. New Phytologist 198, 156-168. C

DeLuca, T.H., Zewdie, S.A., Zackrisson, O., Healey, J.R. & Jones, D.L. (2013). Bracken fern (Pteridium aquilinum L. Kuhn) promotes an open nitrogen cycle in heathland soils. Plant and Soil 367, 521-534. DOI: 10.1007/s11104-012-1484-0.


Tree disease bioeconomic modelling

Macpherson, M.F., Kleczkowski, A., Healey, J.R., Quine, C. & Hanley, N. (2017). The effects of invasive pests and diseases on strategies for forest diversification. Ecological Modelling 350, 87-99.

Macpherson, M.F., Kleczkowski, A., Healey, J.R. & Hanley, N. (2016). The effects of disease on optimal forest rotation: a generalisable analytical framework. Environmental and Resource Economics (published on-line,

Macpherson, M.F., Kleczkowski, A., Healey, J.R. & Hanley, N. (2017). Payment for multiple forest benefits alters the effect of tree disease on optimal forest rotation length. Ecological Economics 134, 82-94.

Sheremet, O., Healey, J.R., Quine, C. & Hanley, N. (2017). Public preferences and willingness to pay for forest disease control in the UK. Journal of Agricultural Economics (published on-line)


Nutrient cycling/resource use/sustainable production

Withers, P.J.A., Sylvester-Bradley, R., Jones, D.L., Healey, J.R. & Talboys, P.J. (2014). Feed the crop not the soil: rethinking phosphorus management in the food chain. Environmental Science & Technology 48, 6523-6530.

Talboys, P.J., Heppell, J., Roose, T., Healey, J.R., Jones, D.L. & Withers, P.J.A. (2016). Struvite: a slow-release fertiliser for sustainable phosphorus management? Plant and Soil 401, 109-123.  DOI:10.1007/s11104-015-2747-3.

Talboys, P.J., Healey, J.R., Withers, P.J.A. & Jones, D.L. (2014). Phosphate depletion modulates auxin transport in Triticum aestivium leading to altered root branching. Journal of Experimental Botany 65, 5023-5032.  DOI:10.1093/jxb/eru284.

Talboys, P.J., Owen, D.W., Healey, J.R., Withers, P.J.A. & Jones, D.L. (2014). Auxin secretion by Bacillus amyloliquefaciens FZB42 both stimulates root exudation and limits phosphorus uptake in Triticum aestivium. BMC Plant Biology 14: 51.  DOI:10.1186/1471-2229-14-51 (

Styles, D., Gibbons, J., Williams, A.P., Stichnothe, H., Chadwick, D.R. & Healey, J.R. (2015). Cattle feed or bioenergy? Consequential life cycle assessment of biogas feedstock options on dairy farms. Global Change Biology Bioenergy 7, 1034-1049.  DOI: 10.1111/gcbb.12189. C

Gibbons, J.M., Williamson, J.C., Williams, A.P., Withers, P.J.A., Hockley, N., Harris, I.M., Hughes, J.W., Taylor, R.L., Jones, D.L. & Healey, J.R. (2014). Sustainable nutrient management at field, farm and regional level: soil testing, nutrient budgets and the trade-off between lime application and greenhouse gas emissions. Agriculture, Ecosystems & Environment 188, 48-56. C

Ford, H., Smith, A., Pagella, T. & Healey, J. (2016). Trees, water storage and flooding in upland agricultural landscapes: why do we need to know more? Forestry & Timber News April 2016, 27-28.


Science-policy interface

O’Leary, B.C., Kvist, K., Bayliss, H.R., Derroire, G., Healey, J.R., Hughes, K., Kleinschroth, F., Sciberras, M., Woodcock, P. & Pullin, A.S. (2016). The reliability of evidence reviews in environmental science and conservation. Environmental Science and Policy 64, 75-82 (




Responsibilities within BU

College of Natural Sciences, Director of Research


Qualifications and Membership of Professional Bodies

BA Botany (University of Oxford)
PhD (University of Cambridge)
Member of the Institute of Chartered Foresters

ID: 4785687