Dr Craig Shuttleworth
Honorary Visiting Research Fellow

Dolenni cyswllt
Contact info
Mobile 07966150847
Email c.shuttleworth@bangor.ac.uk
- 1995
- Cyhoeddwyd
Red squirrels (Sciurus vulgaris) on the Sefton Coast: The conservation value of nest boxes
Shuttleworth, C., 1995, 2nd NPI Red Alert UK Forum for Red Squirrel Conservation. Hughes, D. G. & Tew, T. (gol.). The Federation of Zoological Gardens of Great Britain and Ireland, t. 25-32Allbwn ymchwil: Pennod mewn Llyfr/Adroddiad/Trafodion Cynhadledd › Pennod › adolygiad gan gymheiriaid
- 1997
- Cyhoeddwyd
The effect of supplemental feeding on the diet, population density and reproduction of red squirrels
Shuttleworth, C., 1997, The Conservation of Red Squirrels (Sciurus vulgaris L.). Gurnell, J. & Lurz, P. W. W. (gol.). Peoples Trust for Endangered Species, t. 13-24Allbwn ymchwil: Pennod mewn Llyfr/Adroddiad/Trafodion Cynhadledd › Pennod › adolygiad gan gymheiriaid
- Cyhoeddwyd
The impact of woodland management upon the red squirrel population in Ainsdale National Nature Reserve
Shuttleworth, C., 1997Allbwn ymchwil: Llyfr/Adroddiad › Adroddiad Comisiwn
- 1999
- Cyhoeddwyd
The use of nest boxes by the red squirrel Sciurus vulgaris in a coniferous habitat
Shuttleworth, C., 1 Maw 1999, Yn: Mammal Review. 29, t. 61-66Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- 2000
- Cyhoeddwyd
The foraging behaviour and diet of red squirrels (Sciurus vulgaris) receiving supplemental feeding
Shuttleworth, C., 1 Medi 2000, Yn: Wildlife Biology. 6, t. 149-156Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- 2001
- Cyhoeddwyd
Grey squirrel control as a tactic in the conservation of the red squirrel: a positive outcome
Shuttleworth, C., 2001, Advances in Vertebrate Pest Management . Feare, C. J. & Cowan, D. P. (gol.). Filander Verlag, Cyfrol 3. t. 151-168Allbwn ymchwil: Pennod mewn Llyfr/Adroddiad/Trafodion Cynhadledd › Pennod › adolygiad gan gymheiriaid
- Cyhoeddwyd
The management of coastal sand dune woodland for red squirrels (Sciurus vulgaris L.)
Shuttleworth, C. & Gurnell, J., 2001, Coastal Dune Management: Shared Experience of European Conservation Practice. Houston, J., Edmondson, A. S. E. & Rooney, P. J. (gol.). Liverpool University Press, t. 117-127Allbwn ymchwil: Pennod mewn Llyfr/Adroddiad/Trafodion Cynhadledd › Pennod
- Cyhoeddwyd
The use of nest boxes by the red squirrel (Sciurus vulgaris) in conifer plantations
Shuttleworth, C., 2001, Yn: Quarterly Journal of Forestry. 95, t. 225-228Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- Cyhoeddwyd
Traffic related mortality in a red squirrel (Sciurus vulgaris) population receiving supplemental feeding
Shuttleworth, C., 1 Meh 2001, Yn: Urban Ecosystems. 2, t. 109-118Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- Cyhoeddwyd
Interactions between the red squirrel (Sciurus vulgaris), great tit (Parus Major) and jackdaw (Corvus monedula) whilst using nest boxes.
Shuttleworth, C., 1 Hyd 2001, Yn: Journal of Zoology. 255, t. 269-272Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- 2002
- Cyhoeddwyd
Red squirrel conservation on Anglesey, North Wales. A report on the first five years of the Anglesey Red Squirrel Project
Shuttleworth, C., Bailey, M. & Knott, H., 2002Allbwn ymchwil: Llyfr/Adroddiad › Adroddiad Comisiwn
- 2003
- Cyhoeddwyd
Red squirrels on Anglesey
Shuttleworth, C. & Bailey, M., 2003, Yn: Natur Cymru. 6, t. 35-37Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl
- Cyhoeddwyd
A tough nut to crack: red squirrel conservation in Wales
Shuttleworth, C., 1 Hyd 2003, Yn: Biologist. 50, t. 231-235Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- 2004
- Cyhoeddwyd
A study of nest box use by a low density red squirrel (Sciurus vulgaris) population within a commercial conifer plantation
Shuttleworth, C., 2004, Yn: Quarterly Journal of Forestry. 98, t. 113-119Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- 2005
- Cyhoeddwyd
Against a grey tide: What future for the red squirrel?
Shuttleworth, C., 2005, Journal of Practical Ecology & Conservation (Special Series No 4): Living with the Aliens? - Ecology, History, Culture and Management if exotic plants and animals: issues for nature conservation . Rotherham, L. D. (gol.). Cyfrol 4. t. 87-88Allbwn ymchwil: Pennod mewn Llyfr/Adroddiad/Trafodion Cynhadledd › Pennod
- Cyhoeddwyd
Against a grey tide: red squirrel conservation on Anglesey
Shuttleworth, C., 2005, Yn: Quarterly Journal of Forestry. 99, t. 213-214Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- Cyhoeddwyd
Genetic management of the red squirrel, Sciurus vulgaris: a practical approach to regional conservation
Ogden, R., Shuttleworth, C., McEwing, R. & Cesarini, S., 26 Hyd 2005, Yn: Conservation Genetics. 6, t. 511-525Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- 2008
- Cyhoeddwyd
Re-introduction of the red squirrel into Newborough forest on the island of Anglesey, UK
Shuttleworth, C., Kenward, R. E. & Jackson, N., 2008, Global Re-Introduction Perspectives: Re-Introduction Case-Studies from Around the Globe. Soorae, P. S. (gol.). IUCN/SSC Re-introduction Specialist Group, t. 163-166Allbwn ymchwil: Pennod mewn Llyfr/Adroddiad/Trafodion Cynhadledd › Pennod
- Cyhoeddwyd
Adenovirus-associated deaths in red squirrels on Anglesey
Everest, D. J., Shuttleworth, C. & Stidworthy, M. F., 31 Hyd 2008, Yn: The Veterinary record. 163, 14, t. 430Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- 2009
- Cyhoeddwyd
The reintroduction of the red squirrel Sciurus vulgaris to Newborough forest, North Wales: A five year project review
Shuttleworth, C., Kenward, R. E. & Jackson, N., 2009Allbwn ymchwil: Llyfr/Adroddiad › Adroddiad Comisiwn
- Cyhoeddwyd
PCR detection of adenovirus in grey squirrels on Anglesey
Everest, D. J., Grierson, S. S., Stidworthy, M. & Shuttleworth, C., 31 Hyd 2009, Yn: Veterinary Record. 165, t. 482Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- 2010
- Cyhoeddwyd
Turning the grey tide: progress in red squirrel recovery
Shuttleworth, C., 1 Meh 2010, Yn: Ecos. 31, t. 27-35Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl
-
Retrospective detection by negative contrast electron microscopy of faecal viral particles in wild red squirrels (Sciurus vulgaris) with suspected enteropathy in Great Britain
Everest, D. J., Stidworthy, M. F., Milne, E. M., Meredith, A. L., Chantery, J., Shuttleworth, C., Blackett, T., Butler, H., Wilkinson, M. & Sainsbury, A. W., 31 Rhag 2010, Yn: Veterinary Record. 167, t. 1007-1010Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- 2012
- Cyhoeddwyd
Red squirrel population monitoring: Clocaenog forest
Shuttleworth, C. & Donoyou, T., 2012, IFCW108/001/002 gol.Allbwn ymchwil: Llyfr/Adroddiad › Adroddiad Comisiwn
- Cyhoeddwyd
A systematic assessment of the impact of adenovirus infection on a captive re-introduction project for red squirrels (Sciurus vulgaris)
Everest, D. J., Shuttleworth, C., Grierson, S. S., Duff, J. P., Jackson, N., Litherland, P. & Kenward, R. E., 31 Awst 2012, Yn: Veterinary Record. 171, t. 176-184Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- Cyhoeddwyd
Integrating red squirrel (Sciurus vulgaris) habitat requirements with the management of pathogenic tree disease in commercial forests in the UK
Shuttleworth, C., Lurz, P. W. W., Geddes, N. & Brown, J., 1 Medi 2012, Yn: Forest Ecology and Management. 279, t. 167-175Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- 2013
- Cyhoeddwyd
Adenovirus infection in red squirrels in areas free from grey squirrels
Everest, D. J., Butler, H., Simpson, V. R., Blackett, T. & Shuttleworth, C., 31 Awst 2013, Yn: The Veterinary record. 173, 8, t. 199-200Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl
- 2014
- Cyhoeddwyd
Red squirrel conservation and disease - Adenovirus in Wales
Shuttleworth, C., 2014, Yn: Natur Cymru. 52, t. 41-44Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl
- Cyhoeddwyd
The role of spruce in red squirrel (Sciurus vulgaris) conservation in the UK
Shuttleworth, C. & Lurz, P. W. W., 2014, Forest Ecosystems: Biodiversity, Management and Conservation. Roberts, N. C. (gol.). Nova Science Publishers, t. 63-78Allbwn ymchwil: Pennod mewn Llyfr/Adroddiad/Trafodion Cynhadledd › Pennod › adolygiad gan gymheiriaid
- Cyhoeddwyd
Inter-specific viral infections: Can the management of captive red squirrel collections help inform scientific research?
Shuttleworth, C., Everest, D. J., McInnes, C. J., Greenwood, A., Jackson, N., Rushton, S. P. & Kenward, R. E., 16 Meh 2014, Yn: Hystrix-Italian Journal of Mammalogy. 25, 1, t. 18-24Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- Cyhoeddwyd
Adenovirus infection: An emerging factor in red squirrel conservation programmes
Everest, D. J., Shuttleworth, C., Stidworthy, M. F., Grierson, S. S., Duff, P. J. & Kenward, R. E., 1 Hyd 2014, Yn: Mammal Review. 44, 3-4, t. 225-233Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- Cyhoeddwyd
Landscape scale impacts of culling upon a European grey squirrel population: can trapping reduce population size and decrease the threat of squirrel pox virus infection for the native red squirrel?
Schuchert, P., Shuttleworth, C., McInnes, C. J., Everest, D. J. & Rushton, S. P., 1 Tach 2014, Yn: Biological Invasions. 16, t. 2381-2391Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- Cyhoeddwyd
Are nest boxes a useful tool in regional red squirrel conservation programs?
Shuttleworth, C. & Schuchert , P., 15 Rhag 2014, Yn: Hystrix-Italian Journal of Mammalogy. 25, t. 91-94Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- 2015
- Cyhoeddwyd
British red squirrel strongholds: challenges for conservation
Gurnell, J., Blackett, T., Butler, H., Lurz, P. W. W., Magris, L. & Shuttleworth, C., 2015, Red squirrels: Ecology, Conservation & Management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Hayward, M. W. (gol.). European Squirrel Initiative, t. 211-232Allbwn ymchwil: Pennod mewn Llyfr/Adroddiad/Trafodion Cynhadledd › Pennod › adolygiad gan gymheiriaid
- Cyhoeddwyd
Developing integrated and applied red squirrel conservation programmes: What lessons can Europe learn from a regional grey squirrel eradication programme in North Wales?
Shuttleworth, C., Schuchert, P., Everest, D. J., McInnes, C. J., Rushton, S. P., Jackson, N. & Kenward, R. E., 2015, Red squirrels: Ecology, Conservation & Management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Hayward, M. W. (gol.). European Squirrel Initiative, t. 233-250Allbwn ymchwil: Pennod mewn Llyfr/Adroddiad/Trafodion Cynhadledd › Pennod › adolygiad gan gymheiriaid
- Cyhoeddwyd
Developing red squirrel re-introduction techniques for use during regional grey squirrel eradication programmes in Europe
Shuttleworth, C., Jackson, N. & Kenward, R. E., 2015, 2015 Global Reintroduction Perspectives. Soorae, P. (gol.). IUCN/SSC Re-introduction Specialist Group, t. 67-74Allbwn ymchwil: Pennod mewn Llyfr/Adroddiad/Trafodion Cynhadledd › Pennod
- Cyhoeddwyd
Future red squirrel conservation in Clocaenog forest
Shuttleworth, C. & Hayward, M., 2015, P21018-0026 gol.Allbwn ymchwil: Llyfr/Adroddiad › Adroddiad Comisiwn
- Cyhoeddwyd
Island Haven: Anglesey red squirrel conservation
Shuttleworth, C., 2015, Shared Experience of Red Squirrel Conservation Practice . Shuttleworth, C. M., Lurz, P. W. W. & Halliwell, E. C. (gol.). European Squirrel Initiative, t. 151-158Allbwn ymchwil: Pennod mewn Llyfr/Adroddiad/Trafodion Cynhadledd › Pennod
- Cyhoeddwyd
Painting the Town Red: Gwynedd red squirrel conservation
Shuttleworth, C., 2015, Shared Experience of Red Squirrel Conservation Practice . Shuttleworth, C. M., Lurz, P. W. W. & Halliwell, E. C. (gol.). European Squirrel Initiative, t. 159-166Allbwn ymchwil: Pennod mewn Llyfr/Adroddiad/Trafodion Cynhadledd › Pennod
- Cyhoeddwyd
Red squirrels: ecology, conservation & management in Europe
Shuttleworth, C., Hayward, M. & Lurz, P. W. W., 2015, European Squirrel Initiative. 330 t.Allbwn ymchwil: Llyfr/Adroddiad › Llyfr › adolygiad gan gymheiriaid
- Cyhoeddwyd
Reintroductions and translocations of red squirrels within Europe
Lawton, C., Waters, C. & Shuttleworth, C., 2015, Red squirrels: Ecology, Conservation & Management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Hayward, M. W. (gol.). European Squirrel Initiative, t. 193-210Allbwn ymchwil: Pennod mewn Llyfr/Adroddiad/Trafodion Cynhadledd › Pennod › adolygiad gan gymheiriaid
- Cyhoeddwyd
Shared experience of red squirrel conservation practice
Shuttleworth, C., Lurz, P. W. W. & Halliwell, E. C., 2015, European Squirrel Initiative. 220 t.Allbwn ymchwil: Llyfr/Adroddiad › Llyfr
- Cyhoeddwyd
Striving for success: an evaluation of local action to conserve red squirrels (Sciurus vulgaris) in Wales
Halliwell, E. C., Shuttleworth, C., Wilberforce, L., Denman, H., Lloyd, I. & Cartmel, S., 2015, Red squirrels: Ecology, Conservation & Management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Hayward, M. W. (gol.). European Squirrel Initiative, t. 175-192Allbwn ymchwil: Pennod mewn Llyfr/Adroddiad/Trafodion Cynhadledd › Pennod › adolygiad gan gymheiriaid
- Cyhoeddwyd
The impact of adenoviruses on red squirrel (Sciurus vulgaris) conservation – with notes on other enteric viruses of potential significance
Everest, D. J., Shuttleworth, C., Grierson, S. S., Dastjerdi, A., Stidworthy, M. F., Duff, J. P. & Sainsbury, A. W., 2015, Red squirrels: Ecology, Conservation & Management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Hayward, M. W. (gol.). European Squirrel Initiative, t. 129-146Allbwn ymchwil: Pennod mewn Llyfr/Adroddiad/Trafodion Cynhadledd › Pennod › adolygiad gan gymheiriaid
- Cyhoeddwyd
Adenovirus detections in juvenile squirrels
Everest, D. J., Dastjerdi, A., Cowan, D., Gomm, M., Shuttleworth, C., McInnes, C. J., Deane, D., Coulter, L., Mill, A., Rushton, S. P., Jackson, N. L. & Litherland, P., 1 Hyd 2015, Yn: Veterinary Record. 177, t. 373-374Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl
- Cyhoeddwyd
Assessing causes and significance of red squirrel (Sciurus vulgaris) mortality during regional population restoration: An applied conservation perspective
Shuttleworth, C., Signorile, , A. L., Everest, D. J., Lurz, P. W. W. & Duff, J. P., 16 Tach 2015, Yn: Hystrix-Italian Journal of Mammalogy. 26, 2, t. 69-75Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- 2016
- Cyhoeddwyd
Ecosystem impacts of an alien invader in Europe, the grey squirrel Sciurus carolinensis
Gurnell, J., Lurz, P. W. W. & Shuttleworth, C., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (gol.). European Squirrel Initiative, t. 307-329Allbwn ymchwil: Pennod mewn Llyfr/Adroddiad/Trafodion Cynhadledd › Pennod › adolygiad gan gymheiriaid
- Cyhoeddwyd
Experiences of grey squirrel management in an upland conifer forest
Halliwell, E. C., Shuttleworth, C., Cartmel, S., Lloyd, I. & Jenkins, R., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (gol.). European Squirrel Initiative, t. 453-474Allbwn ymchwil: Pennod mewn Llyfr/Adroddiad/Trafodion Cynhadledd › Pennod › adolygiad gan gymheiriaid
- Cyhoeddwyd
Genetic evidence of the effectiveness of grey squirrel control operations: lessons from the Isle of Anglesey
Signorile, L. & Shuttleworth, C., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (gol.). European Squirrel Initiative, t. 439-452Allbwn ymchwil: Pennod mewn Llyfr/Adroddiad/Trafodion Cynhadledd › Pennod › adolygiad gan gymheiriaid
- Cyhoeddwyd
Grey squirrel management in Europe - the future
Shuttleworth, C., Lurz, P. W. W. & Gurnell, J., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (gol.). European Squirrel InitiativeAllbwn ymchwil: Pennod mewn Llyfr/Adroddiad/Trafodion Cynhadledd › Pennod › adolygiad gan gymheiriaid