Dr Craig Shuttleworth
Honorary Visiting Research Fellow

Dolenni cyswllt
Contact info
Mobile 07966150847
Email c.shuttleworth@bangor.ac.uk
- Cyhoeddwyd
A practical guide to early surveillance and monitoring
Robinson, N. J., Everest, D. J., Harry, P. & Shuttleworth, C., 2021, Invasive Alien Species Colonisation Prevention: Your guide to early detection and rapid response . Robinson, N. J. & Shuttleworth, C. M. (gol.). Royal Society of Wildlife Trusts, t. 163-202Allbwn ymchwil: Pennod mewn Llyfr/Adroddiad/Trafodion Cynhadledd › Pennod
- Cyhoeddwyd
A preliminary comparison of magnum 116 body-grip traps with a live capture trap design during the control of grey squirrels (Sciurus carolinensis)
Shuttleworth, C., Mill, A. & Van der Waal, Z., Rhag 2016, Yn: International Journal of Pest Management. 63, 4, t. 316-321Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- Cyhoeddwyd
A study of nest box use by a low density red squirrel (Sciurus vulgaris) population within a commercial conifer plantation
Shuttleworth, C., 2004, Yn: Quarterly Journal of Forestry. 98, t. 113-119Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- Cyhoeddwyd
A systematic assessment of the impact of adenovirus infection on a captive re-introduction project for red squirrels (Sciurus vulgaris)
Everest, D. J., Shuttleworth, C., Grierson, S. S., Duff, J. P., Jackson, N., Litherland, P. & Kenward, R. E., 31 Awst 2012, Yn: Veterinary Record. 171, t. 176-184Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- Cyhoeddwyd
A temporal refuge from predation can change the outcome of prey species competition
Slade, A., White, A., Lurz, P. W. W., Shuttleworth, C. & Lambin, X., 1 Medi 2022, Yn: Oikos. Oikos e08565Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- Cyhoeddwyd
A tough nut to crack: red squirrel conservation in Wales
Shuttleworth, C., 1 Hyd 2003, Yn: Biologist. 50, t. 231-235Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- Cyhoeddwyd
Adenovirus detections in juvenile squirrels
Everest, D. J., Dastjerdi, A., Cowan, D., Gomm, M., Shuttleworth, C., McInnes, C. J., Deane, D., Coulter, L., Mill, A., Rushton, S. P., Jackson, N. L. & Litherland, P., 1 Hyd 2015, Yn: Veterinary Record. 177, t. 373-374Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl
- Cyhoeddwyd
Adenovirus infection in red squirrels
Shuttleworth, C., Everest, D. J., Stidworthy, M., Halliwell, E. C., Clews-Roberts, R., Paine, V., Bamber, C. & Mason, J., 31 Rhag 2021, Yn: The Veterinary record. 189, t. 444-445Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl
- Cyhoeddwyd
Adenovirus infection in red squirrels in areas free from grey squirrels
Everest, D. J., Butler, H., Simpson, V. R., Blackett, T. & Shuttleworth, C., 31 Awst 2013, Yn: The Veterinary record. 173, 8, t. 199-200Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl
- Cyhoeddwyd
Adenovirus infection: An emerging factor in red squirrel conservation programmes
Everest, D. J., Shuttleworth, C., Stidworthy, M. F., Grierson, S. S., Duff, P. J. & Kenward, R. E., 1 Hyd 2014, Yn: Mammal Review. 44, 3-4, t. 225-233Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- Cyhoeddwyd
Adenovirus remains widespread in squirrels on Anglesey
Shuttleworth, C., Everest, D. & McQueen, A., 16 Rhag 2023, Yn: Veterinary Record. 193, 12, t. 490-491 2 t.Allbwn ymchwil: Cyfraniad at gyfnodolyn › Llythyr
- Cyhoeddwyd
Adenovirus-associated deaths in red squirrels on Anglesey
Everest, D. J., Shuttleworth, C. & Stidworthy, M. F., 31 Hyd 2008, Yn: The Veterinary record. 163, 14, t. 430Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- Cyhoeddwyd
Against a grey tide: What future for the red squirrel?
Shuttleworth, C., 2005, Journal of Practical Ecology & Conservation (Special Series No 4): Living with the Aliens? - Ecology, History, Culture and Management if exotic plants and animals: issues for nature conservation . Rotherham, L. D. (gol.). Cyfrol 4. t. 87-88Allbwn ymchwil: Pennod mewn Llyfr/Adroddiad/Trafodion Cynhadledd › Pennod
- Cyhoeddwyd
Against a grey tide: red squirrel conservation on Anglesey
Shuttleworth, C., 2005, Yn: Quarterly Journal of Forestry. 99, t. 213-214Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- Cyhoeddwyd
An Opportunistic Assessment of the Impact of Squirrelpox Disease Outbreaks upon a Red Squirrel Population Sympatric with Grey Squirrels in Wales
Shuttleworth, C., Everest, D., Holmes, P., Bell, S. & Cripps, R., 1 Ion 2022, Yn: Animals. 12, 1Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- Cyhoeddwyd
Are nest boxes a useful tool in regional red squirrel conservation programs?
Shuttleworth, C. & Schuchert , P., 15 Rhag 2014, Yn: Hystrix-Italian Journal of Mammalogy. 25, t. 91-94Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- Cyhoeddwyd
Are novel ecosystems the only novelty of rewilding?
Klop-Toker, K., Clulow, S., Shuttleworth, C. & Hayward, M., 1 Tach 2020, Yn: Restoration Ecology. 28, t. 1318-1320Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- Cyhoeddwyd
Assessing a potential non-invasive method for viral diagnostic purposes in European squirrels
Everest, D. J., Tolhurst-Cherriman, D. A. R., Davies, H., Dastjerdi, A., Ashton, A., Blackett, T., Meredith, A. L., Milne, E., Mill, A. & Shuttleworth, C., 4 Meh 2019, Yn: Hystrix-Italian Journal of Mammalogy. 30, t. 44-50Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- Cyhoeddwyd
Assessing causes and significance of red squirrel (Sciurus vulgaris) mortality during regional population restoration: An applied conservation perspective
Shuttleworth, C., Signorile, , A. L., Everest, D. J., Lurz, P. W. W. & Duff, J. P., 16 Tach 2015, Yn: Hystrix-Italian Journal of Mammalogy. 26, 2, t. 69-75Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- Cyhoeddwyd
British red squirrel strongholds: challenges for conservation
Gurnell, J., Blackett, T., Butler, H., Lurz, P. W. W., Magris, L. & Shuttleworth, C., 2015, Red squirrels: Ecology, Conservation & Management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Hayward, M. W. (gol.). European Squirrel Initiative, t. 211-232Allbwn ymchwil: Pennod mewn Llyfr/Adroddiad/Trafodion Cynhadledd › Pennod › adolygiad gan gymheiriaid
- Cyhoeddwyd
British red squirrels remain the only known wild rodent host for leprosy bacilli
Schilling, A., Avanzi, C., Ulrich, R. G., Busso, P., Pisanu, B., Ferrari, N., Romeo, C., Mazzamuto, M. V., McLuckie, J., Shuttleworth, C., Del-Pozo, J., Lurz, P. W. W., Escalante-Fuentes, W. G., Ocampo-Candiani, J., Vera-Cabrera, L., Stevenson, K., Chapuis, J.-L., Meredith, A. L. & Cole, S. T., 1 Chwef 2019, Yn: Frontiers in Veterinary Science. 6, t. 8Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- Cyhoeddwyd
By trying to be everything, rewilding risks meaning nothing
Jachowski, J. D., Bombaci, S., Caravaggi, A., Chalmers, A., Clulow, S., Griffin, A., Hayward, M., Heurich, M., Linnell, J., Marnewick, K., Moehrenschlager, A., Montgomery, A., Shuttleworth, C., Somers, M. & Weise, F., Tach 2019, Yn: Science. 364, t. 6438Allbwn ymchwil: Cyfraniad at gyfnodolyn › Llythyr
- Cyhoeddwyd
Capturing red squirrels (Sciurus vulgaris) on camera: A cost-effective approach for monitoring relative abundance and habitat preference
Shannon, G., Valle, S. & Shuttleworth, C., 3 Hyd 2023, Yn: Ecology and Evolution. 13, 10, e10536.Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- Cyhoeddwyd
Challenges and solutions in early detection, rapid response and communication about potential invasive alien species in forests
de Groot, M., Bullas-Appleton, E., Csóka, G., Csiszár, A., Faccoli, A., Gervasini, A., Kirichenko, N., Shuttleworth, C., Kus Veenvliet, J., O'Hanlon, R., Korda, M., Marinsek, A., Robinson, N., Sweeney, J., Tricarico, E., Verbrugge, L., Williams, D. & Zidar, S., 14 Awst 2020, Yn: Management of Biological Invasions. 11, t. 637-660Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- Cyhoeddwyd
Confirmation of squirrelpox in Welsh red squirrel
Everest, D. J., Floyd, T., Donnnacie, B., Irvine, R. M., Holmes, J. P. & Shuttleworth, C., 30 Tach 2017, Yn: Veterinary Record. 181, t. 514-515Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl
- Cyhoeddwyd
Controlling grey squirrels in forests and woodlands in the UK
Gill, R., Ferryman, M., Shuttleworth, C., Lurz, P. W. W., Mill, A., Robertson, P. A. & Dutton, C., 2019, Forest Research. 16 t.Allbwn ymchwil: Llyfr/Adroddiad › Llyfr › adolygiad gan gymheiriaid
- Cyhoeddwyd
Deconstructing compassionate conservation
Hayward, M. W., Callen, A., Allen, B. L., Ballard, G., Broekhuis, F., Bugir, C., Clarke, R. H., Clulow, J., Clulow, S., Daltry, J. C., Davies-Mostert, H. T., Fleming, P. J. S., Griffin, A. S., Howell, L. G., Kerley, G. I. H., Klop-Toker, K., Legge, S., Major, T., Meyer, N., Montgomery, R. A., Moseby, K., Parker, D. M., Périquet, S., Read, J., Scanlon, R., Seeto, R., Shuttleworth, C., Somers, M. J., Tamessar, C. T., Tuft, K., Upton, R., Valenzuela-Molina, M., Wayne, A., Witt, R. R. & Wuster, W., Awst 2019, Yn: Conservation Biology. 33, 4, t. 760-768Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- Cyhoeddwyd
Detecting viral infection in red squirrels
Shuttleworth, C., Everest, D. J., Halliwell, E. C., Hulme, B., Wilberforce, L. & Clews-Roberts, R., 30 Ebr 2019, Yn: Veterinary Record. 184, t. 507Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl
- Cyhoeddwyd
Developing a method for assessing the relative abundance of red squirrels in low-density populations
Shannon, G., Barton, O., Valle, S. & Shuttleworth, C., 1 Tach 2024, 34 t.Allbwn ymchwil: Llyfr/Adroddiad › Adroddiad Comisiwn
- Cyhoeddwyd
Developing integrated and applied red squirrel conservation programmes: What lessons can Europe learn from a regional grey squirrel eradication programme in North Wales?
Shuttleworth, C., Schuchert, P., Everest, D. J., McInnes, C. J., Rushton, S. P., Jackson, N. & Kenward, R. E., 2015, Red squirrels: Ecology, Conservation & Management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Hayward, M. W. (gol.). European Squirrel Initiative, t. 233-250Allbwn ymchwil: Pennod mewn Llyfr/Adroddiad/Trafodion Cynhadledd › Pennod › adolygiad gan gymheiriaid
- Cyhoeddwyd
Developing red squirrel re-introduction techniques for use during regional grey squirrel eradication programmes in Europe
Shuttleworth, C., Jackson, N. & Kenward, R. E., 2015, 2015 Global Reintroduction Perspectives. Soorae, P. (gol.). IUCN/SSC Re-introduction Specialist Group, t. 67-74Allbwn ymchwil: Pennod mewn Llyfr/Adroddiad/Trafodion Cynhadledd › Pennod
- Cyhoeddwyd
Disease monitoring and surveillance: case studies in the applied conservation of fragmented red squirrel (Sciurus vulgaris) populations in England and Wales
Everest, D. J., Floyd, T., Holmes, P., Man, C., Dunnett, C., Locke, R., Savage, L., Sutcliffe, S., Sapsford, B., Shuttleworth, C. & Duff, P., 1 Rhag 2021, Yn: Mammalian Biology. 101, t. 1079-1088Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- Cyhoeddwyd
Disease, invasions and conservation: no evidence of squirrelpox virus in grey squirrels introduced to Italy
Romeo, C., McInnes, C., Dale, T., Shuttleworth, C., Bertolino, S., Wauters, L. A. & Ferrari, N., 1 Chwef 2019, Yn: Animal Conservation. 22, 1, t. 14-23Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- Cyhoeddwyd
Do differing levels of boldness influence the success of translocation? A pilot study on red squirrels (Sciurus vulgaris)
Bamber, J. A., Shuttleworth, C. & Hayward, M., 26 Medi 2020, Yn: Animals. 10, 1748.Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- Cyhoeddwyd
Ecosystem impacts of an alien invader in Europe, the grey squirrel Sciurus carolinensis
Gurnell, J., Lurz, P. W. W. & Shuttleworth, C., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (gol.). European Squirrel Initiative, t. 307-329Allbwn ymchwil: Pennod mewn Llyfr/Adroddiad/Trafodion Cynhadledd › Pennod › adolygiad gan gymheiriaid
- Cyhoeddwyd
Envisioning the future with 'compassionate conservation': An ominous projection for native wildlife and biodiversity
Callen, A., Hayward, M. W., Klop-Toker, K., Allen, B. L., Ballard, G., Beranek, C. T., Broekhuis, F., Bugir, C. K., Clarke, R. H., Clulow, J., Clulow, S., Daltry, J. C., Davies-Mostert, H. T., Di Blanco, Y. E., Dixon, V., Fleming, P. J. S., Howell, L. G., Kerley, G. I. H., Legge, S. M., Lenga, D. J., Major, T., Montgomery, R. A., Moseby, K., Meyer, N., Parker, D. M., Periquet, S., Read, J., Scanlon, R. J., Shuttleworth, C., Tamessar, C. T., Taylor, W. A., Tuft, K., Upton, R. M. O., Valenzuela, M., Witt, R. R. & Wuster, W., Ion 2020, Yn: Biological Conservation. 241, 108365.Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
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Ethical dilemmas when using citizen science for early detection of invasive tree pests and diseases
Pocock, M. J. O., Marzano, M., Bullas-Appleton, E., Dyke, A., de Groot, M., Shuttleworth, C. & White, R., 1 Tach 2020, Yn: Management of Biological Invasions. 11, t. 720-732Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- Cyhoeddwyd
Evolving grey squirrel management techniques in Europe
Shuttleworth, C., Robinson, N. J., Halliwell, E. C., Clews-Roberts, R., Peek, H., Podgornik, G., Stinson, M., Rice, S., Finlay, C., McKinney, C., Everest, D. J. & Larsen, K. W., 21 Hyd 2020, Yn: Management of Biological Invasions. 11, 4, t. 747-761Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- Cyhoeddwyd
Experiences of grey squirrel management in an upland conifer forest
Halliwell, E. C., Shuttleworth, C., Cartmel, S., Lloyd, I. & Jenkins, R., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (gol.). European Squirrel Initiative, t. 453-474Allbwn ymchwil: Pennod mewn Llyfr/Adroddiad/Trafodion Cynhadledd › Pennod › adolygiad gan gymheiriaid
- Cyhoeddwyd
First report of dermatophilosis in wild European red squirrels (Sciurus vulgaris)
Holmes, P., Everest, D. J., Spiro, S., Wessels, M. & Shuttleworth, C., 30 Medi 2019, Yn: Veterinary Record Case Reports. 7, 3, t. 3Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- Cyhoeddwyd
Future red squirrel conservation in Clocaenog forest
Shuttleworth, C. & Hayward, M., 2015, P21018-0026 gol.Allbwn ymchwil: Llyfr/Adroddiad › Adroddiad Comisiwn
- Cyhoeddwyd
Genetic evidence of the effectiveness of grey squirrel control operations: lessons from the Isle of Anglesey
Signorile, L. & Shuttleworth, C., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (gol.). European Squirrel Initiative, t. 439-452Allbwn ymchwil: Pennod mewn Llyfr/Adroddiad/Trafodion Cynhadledd › Pennod › adolygiad gan gymheiriaid
- Cyhoeddwyd
Genetic management of the red squirrel, Sciurus vulgaris: a practical approach to regional conservation
Ogden, R., Shuttleworth, C., McEwing, R. & Cesarini, S., 26 Hyd 2005, Yn: Conservation Genetics. 6, t. 511-525Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- Cyhoeddwyd
Grey squirrel control as a tactic in the conservation of the red squirrel: a positive outcome
Shuttleworth, C., 2001, Advances in Vertebrate Pest Management . Feare, C. J. & Cowan, D. P. (gol.). Filander Verlag, Cyfrol 3. t. 151-168Allbwn ymchwil: Pennod mewn Llyfr/Adroddiad/Trafodion Cynhadledd › Pennod › adolygiad gan gymheiriaid
- Cyhoeddwyd
Grey squirrel management in Europe - the future
Shuttleworth, C., Lurz, P. W. W. & Gurnell, J., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (gol.). European Squirrel InitiativeAllbwn ymchwil: Pennod mewn Llyfr/Adroddiad/Trafodion Cynhadledd › Pennod › adolygiad gan gymheiriaid
- Cyhoeddwyd
Grey squirrel nesting ecology and the use of nest sites in European population management
Shuttleworth, C., Selonen, V. & Koprowski, J. L., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (gol.). European Squirrel Initiative, t. 349-369Allbwn ymchwil: Pennod mewn Llyfr/Adroddiad/Trafodion Cynhadledd › Pennod › adolygiad gan gymheiriaid
- Cyhoeddwyd
Grey squirrels are bad for the British countryside – full stop
Shuttleworth, C., 2017, The Conversation.Allbwn ymchwil: Cyfraniad arall › Cyfraniad Arall
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Gut bacteria may explain why grey squirrels outcompete reds – new research
Shuttleworth, C., 20 Chwef 2024, The Conversation.Allbwn ymchwil: Cyfraniad at gyhoeddiad arbenigol › Erthygl
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Identifying incursion pathways, early detection responses and management actions to prevent grey squirrel range expansion: an island case study in Wales
Shuttleworth, C., Halliwell, E. C. & Robertson, P. A., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (gol.). European Squirrel Initiative, t. 475-494Allbwn ymchwil: Pennod mewn Llyfr/Adroddiad/Trafodion Cynhadledd › Pennod › adolygiad gan gymheiriaid
- Cyhoeddwyd
Indirect effects of pine marten recovery result in benefits to native prey through suppression of an invasive species and a shared pathogen
Slade, A., White, A., Lurz, P. W. W., Shuttleworth, C., Tosh, D. & Twining, J., 1 Chwef 2023, Yn: Ecological Modelling. 476, 110216.Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid