Dr Craig Shuttleworth
Honorary Visiting Research Fellow

Dolenni cyswllt
Contact info
Mobile 07966150847
Email c.shuttleworth@bangor.ac.uk
- Cyhoeddwyd
Why the pine marten is not every red squirrel’s best friend
Shuttleworth, C. & Hayward, M., 2019, The Conversation.Allbwn ymchwil: Cyfraniad arall › Cyfraniad Arall
- Cyhoeddwyd
Wales should follow Scotland’s lead in protecting the red squirrel’s habitat
Shuttleworth, C., 2018, Nation Cymru .Allbwn ymchwil: Cyfraniad arall › Cyfraniad Arall
- Cyhoeddwyd
Virus presence within Welsh red squirrels
Everest, D. J., Holmes, P. & Shuttleworth, C., 30 Mai 2017, Yn: The Veterinary record. 180, t. 520-521Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl
- Cyhoeddwyd
Turning the grey tide: progress in red squirrel recovery
Shuttleworth, C., 1 Meh 2010, Yn: Ecos. 31, t. 27-35Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl
- Cyhoeddwyd
Traffic related mortality in a red squirrel (Sciurus vulgaris) population receiving supplemental feeding
Shuttleworth, C., 1 Meh 2001, Yn: Urban Ecosystems. 2, t. 109-118Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- Cyhoeddwyd
The use of nest boxes by the red squirrel Sciurus vulgaris in a coniferous habitat
Shuttleworth, C., 1 Maw 1999, Yn: Mammal Review. 29, t. 61-66Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- Cyhoeddwyd
The use of nest boxes by the red squirrel (Sciurus vulgaris) in conifer plantations
Shuttleworth, C., 2001, Yn: Quarterly Journal of Forestry. 95, t. 225-228Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- Cyhoeddwyd
The search for novelty continues for rewilding
Hayward, M., Jachowski, J. D., Bugir, C. J., Clulow, J., Krishnamurthy, R., Griffin, A. S., Chalmers, A. C., Linnell, J. D. C., Montgomery, R. A., Somers, M. J., Kowalczyk, R., Heurich, M., Caravaggi, A., Marnewick, K. A., Di Blanco, Y., Shuttleworth, C., Callen, C., Weise, F., Scanlon, R., Moehrenschlager, A., Howell, L. & Upton, R. M. O., 1 Awst 2019, Yn: Biological Conservation. 236, t. 584-585Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- Cyhoeddwyd
The role of spruce in red squirrel (Sciurus vulgaris) conservation in the UK
Shuttleworth, C. & Lurz, P. W. W., 2014, Forest Ecosystems: Biodiversity, Management and Conservation. Roberts, N. C. (gol.). Nova Science Publishers, t. 63-78Allbwn ymchwil: Pennod mewn Llyfr/Adroddiad/Trafodion Cynhadledd › Pennod › adolygiad gan gymheiriaid
- Cyhoeddwyd
The reintroduction of the red squirrel Sciurus vulgaris to Newborough forest, North Wales: A five year project review
Shuttleworth, C., Kenward, R. E. & Jackson, N., 2009Allbwn ymchwil: Llyfr/Adroddiad › Adroddiad Comisiwn
- Cyhoeddwyd
The potential role of ectoparasites in the epidemiology of squirrelpox virus: a possible novel means of intervention to reduce the impact of the disease?
Cowan, D. P., Mill, A. C., Everest, D. J., McInnes, C. J., Rushton, S. P., Shirley, M. D. F., Start, C. & Shuttleworth, C., 2016, The grey squirrel: ecology and management in Europe . Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (gol.). European Squirrel Initiative, t. 255-289Allbwn ymchwil: Pennod mewn Llyfr/Adroddiad/Trafodion Cynhadledd › Pennod › adolygiad gan gymheiriaid
- Cyhoeddwyd
The management of grey squirrel populations in Europe: evolving best practice
Bertolino, S., Lurz, P. W. W., Shuttleworth, C., Martinoli, A. & Wauters, L. A., 2016, The grey squirrel: ecology and management in Europe . Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (gol.). European Squirrel Initiative, t. 495-516Allbwn ymchwil: Pennod mewn Llyfr/Adroddiad/Trafodion Cynhadledd › Pennod › adolygiad gan gymheiriaid
- Cyhoeddwyd
The management of coastal sand dune woodland for red squirrels (Sciurus vulgaris L.)
Shuttleworth, C. & Gurnell, J., 2001, Coastal Dune Management: Shared Experience of European Conservation Practice. Houston, J., Edmondson, A. S. E. & Rooney, P. J. (gol.). Liverpool University Press, t. 117-127Allbwn ymchwil: Pennod mewn Llyfr/Adroddiad/Trafodion Cynhadledd › Pennod
- Cyhoeddwyd
The large-scale removal of mammalian invasive alien species in Northern Europe
Robertson, P. A., Adriaens, T., Lambin, X., Mill, A., Roy, S., Shuttleworth, C. & Sutton-Croft, M., 1 Chwef 2017, Yn: Pest Management Science. 73, t. 273-279Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- Cyhoeddwyd
The implications of significant adenovirus infection in UK captive red squirrel (Sciurus vulgaris) collections: How histological screening can aid applied conservation management
Everest, D. J., Shuttleworth, C., Grierson, S. S., Dastjerdi, A., M.F., S., Duff, J. P., Higgins, R. J., Mill, A. C. & Chantrey, J., 1 Ion 2018, Yn: Mammalian Biology. 88, t. 123-129Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- Cyhoeddwyd
The impact of woodland management upon the red squirrel population in Ainsdale National Nature Reserve
Shuttleworth, C., 1997Allbwn ymchwil: Llyfr/Adroddiad › Adroddiad Comisiwn
- Cyhoeddwyd
The impact of adenoviruses on red squirrel (Sciurus vulgaris) conservation – with notes on other enteric viruses of potential significance
Everest, D. J., Shuttleworth, C., Grierson, S. S., Dastjerdi, A., Stidworthy, M. F., Duff, J. P. & Sainsbury, A. W., 2015, Red squirrels: Ecology, Conservation & Management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Hayward, M. W. (gol.). European Squirrel Initiative, t. 129-146Allbwn ymchwil: Pennod mewn Llyfr/Adroddiad/Trafodion Cynhadledd › Pennod › adolygiad gan gymheiriaid
- Cyhoeddwyd
The grey squirrel: ecology & management of an Invasive Species in Europe
Shuttleworth, C., Lurz, P. W. W. & Gurnell, J., 2016, European Squirrel Initiative. 505 t.Allbwn ymchwil: Llyfr/Adroddiad › Llyfr › adolygiad gan gymheiriaid
- Cyhoeddwyd
The foraging behaviour and diet of red squirrels (Sciurus vulgaris) receiving supplemental feeding
Shuttleworth, C., 1 Medi 2000, Yn: Wildlife Biology. 6, t. 149-156Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- Cyhoeddwyd
The effect of supplemental feeding on the diet, population density and reproduction of red squirrels
Shuttleworth, C., 1997, The Conservation of Red Squirrels (Sciurus vulgaris L.). Gurnell, J. & Lurz, P. W. W. (gol.). Peoples Trust for Endangered Species, t. 13-24Allbwn ymchwil: Pennod mewn Llyfr/Adroddiad/Trafodion Cynhadledd › Pennod › adolygiad gan gymheiriaid
- Cyhoeddwyd
Sympatric cases of squirrelpox in red and grey squirrels
Shuttleworth, C., Everest, D., Wood, C. & Holmes, J. P., 7 Mai 2021, Yn: The Veterinary record. 188, 9, t. 358Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- Cyhoeddwyd
Striving for success: an evaluation of local action to conserve red squirrels (Sciurus vulgaris) in Wales
Halliwell, E. C., Shuttleworth, C., Wilberforce, L., Denman, H., Lloyd, I. & Cartmel, S., 2015, Red squirrels: Ecology, Conservation & Management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Hayward, M. W. (gol.). European Squirrel Initiative, t. 175-192Allbwn ymchwil: Pennod mewn Llyfr/Adroddiad/Trafodion Cynhadledd › Pennod › adolygiad gan gymheiriaid
- Cyhoeddwyd
Squirrelpox outbreak detected in north Wales – without a vaccine, the disease will keep decimating red squirrels
Shuttleworth, C., 21 Rhag 2022, The Conversation.Allbwn ymchwil: Cyfraniad arall › Cyfraniad Arall
- Cyhoeddwyd
Significant differences in the caecal bacterial microbiota of red and grey squirrels in Britain
Hall, L., Nichols, C., Leng, J., Martelli, F., Shuttleworth, C. & La Ragione, R., 14 Chwef 2024, Yn: Journal of Medical Microbiology. 73, 2Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- Cyhoeddwyd
Shared experience of red squirrel conservation practice
Shuttleworth, C., Lurz, P. W. W. & Halliwell, E. C., 2015, European Squirrel Initiative. 220 t.Allbwn ymchwil: Llyfr/Adroddiad › Llyfr
- Cyhoeddwyd
Saving the red squirrel: landscape scale recovery
Shuttleworth, C., Lurz, P. W. W. & Robinson, N., 2021, London: Red Squirrel Survival Trust. 306 t.Allbwn ymchwil: Llyfr/Adroddiad › Llyfr › adolygiad gan gymheiriaid
- Cyhoeddwyd
Rewilding needs to clarify the role of management of invasive species
Hayward, M., Jachowski, D. S., Shuttleworth, C., Linnell, J. D. C., Allen, B. L., Griffen, A. S., Montgomery, R. A., Caravaggi, A., Weise, F. J., Moehrenschlager, A., Somers, M. J., Clulow, S., Heurich, M. & Marnewick, K., 8 Mai 2019, Yn: Science. 364, t. 6438Allbwn ymchwil: Cyfraniad at gyfnodolyn › Llythyr
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Retrospective detection by negative contrast electron microscopy of faecal viral particles in wild red squirrels (Sciurus vulgaris) with suspected enteropathy in Great Britain
Everest, D. J., Stidworthy, M. F., Milne, E. M., Meredith, A. L., Chantery, J., Shuttleworth, C., Blackett, T., Butler, H., Wilkinson, M. & Sainsbury, A. W., 31 Rhag 2010, Yn: Veterinary Record. 167, t. 1007-1010Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- Cyhoeddwyd
Restoring our lost wildlife: a review of the ecological, social and economic factors in the potential reintroduction of six mammal species to Cornwall
Cooper, P., Brazier, R., Puttock, A., Graham, H., Auster, R., Campbell-Palmer, R., Shuttleworth, C. & Pizzi, R., 2023, 258 t.Allbwn ymchwil: Llyfr/Adroddiad › Adroddiad Comisiwn
- Cyhoeddwyd
Response to comments on "Compassionate Conservation deserves a morally serious rather than dismissive response - reply to Callen et al., 2020"
Callen, A., Hayward, M. W., Klop-Toker, K., Allen, B. L., Ballard, G., Beranek, C. T., Broekhuis, F., Bugir, C. K., Clarke, R. H., Clulow, J., Clulow, S., Daltry, J. C., Davies-Mostert, H. T., Di Blanco, Y. E., Dixon, V., Fleming, P. J. S., Howell, L. G., Kerley, G. I. H., Legge, S. M., Lenga, D. J., Major, T., Montgomery, R. A., Moseby, K., Meyer, N., Parker, D. M., Periquet, S., Read, J., Scanlon, R. J., Shuttleworth, C., Tamessar, C. T., Taylor, W. A., Tuft, K., Upton, R. M. O., Valenzuela, M., Witt, R. R. & Wuster, W., 30 Ebr 2020, Yn: Biological Conservation. 244Allbwn ymchwil: Cyfraniad at gyfnodolyn › Llythyr › adolygiad gan gymheiriaid
- Cyhoeddwyd
Repeated squirrelpox infection in Welsh red squirrels
Holmes, J. P., Everest, D. & Shuttleworth, C., 3 Chwef 2023, Yn: Veterinary Record. 192, 3, t. 128-129Allbwn ymchwil: Cyfraniad at gyfnodolyn › Llythyr
- Cyhoeddwyd
Releasing grey squirrels into the wild
Shuttleworth, C., Bertolino, S., Gill, R., Gurnell, J., Hayward, M., Kenward, R. E., Lawton, C., Lurz, P. W. W., McInnes, C. J., Mill, A., Trotter, S. & Wauters, L. A., 31 Maw 2019, Yn: Veterinary Record. 184, t. 389-390Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl
- Cyhoeddwyd
Reintroductions and translocations of red squirrels within Europe
Lawton, C., Waters, C. & Shuttleworth, C., 2015, Red squirrels: Ecology, Conservation & Management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Hayward, M. W. (gol.). European Squirrel Initiative, t. 193-210Allbwn ymchwil: Pennod mewn Llyfr/Adroddiad/Trafodion Cynhadledd › Pennod › adolygiad gan gymheiriaid
- Cyhoeddwyd
Reintroducing rewilding to restoration – Rejecting the search for novelty
Hayward, M. W., Scanlon, R. J., Callen, A., Howell, L. G., Klop-Toker, K. L., Di Blanco, Y., Balkenhol, N., Bugir, C. K., Campbell, L., Caravaggi, A., Chalmers, A. C., Clulow, J., Clulow, S., Cross, P., Gould, J. A., Griffin, A. S., Heurich, M., Howe, B. K., Jachowski, D. S., Jhala, Y. V., Krishnamurthy, R., Kowalczyk, R., Lenga, D. J., Linnell, J. D. C., Marnewick, K. A., Moehrenschlager, A., Montgomery, R. A., Osipova, L., Peneaux, C., Rodger, J. C., Sales, L. P., Seeto, R. G. Y., Shuttleworth, C. M., Somers, M. J., Tamessar, C. T., Upton, R. M. O. & Weise, F. J., 1 Mai 2019, Yn: Biological Conservation. 233, t. 255-259 5 t.Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- Cyhoeddwyd
Reinstating trophic cascades as an applied conservation tool to protect forest ecosystems from invasive grey squirrels (Sciurus carolinensis)
Bamber, J. A., Shuttleworth, C. M., Hayward, M. W. & Everest, D. J., 31 Rhag 2020, Yn: Food Webs. 25, e00164.Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- Cyhoeddwyd
Reducing negative impacts on non-target species during grey squirrel (Sciurus carolinensis) control activities
Shuttleworth, C., Mill, A. C., Van der Wall, Z., Bertolino, S. & Everest, D. J., 1 Ion 2018, Yn: Quarterly Journal of Forestry. 112, 1, t. 21-29Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- Cyhoeddwyd
Red squirrels: ecology, conservation & management in Europe
Shuttleworth, C., Hayward, M. & Lurz, P. W. W., 2015, European Squirrel Initiative. 330 t.Allbwn ymchwil: Llyfr/Adroddiad › Llyfr › adolygiad gan gymheiriaid
- Cyhoeddwyd
Red squirrels on Anglesey
Shuttleworth, C. & Bailey, M., 2003, Yn: Natur Cymru. 6, t. 35-37Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl
- Cyhoeddwyd
Red squirrels in my garden: guidance and tips to help conserve local populations
Shuttleworth, C. & Halliwell, E. C., 2018, European Squirrel Initiative. 185 t.Allbwn ymchwil: Llyfr/Adroddiad › Llyfr › adolygiad gan gymheiriaid
- Cyhoeddwyd
Red squirrels (Sciurus vulgaris) on the Sefton Coast: The conservation value of nest boxes
Shuttleworth, C., 1995, 2nd NPI Red Alert UK Forum for Red Squirrel Conservation. Hughes, D. G. & Tew, T. (gol.). The Federation of Zoological Gardens of Great Britain and Ireland, t. 25-32Allbwn ymchwil: Pennod mewn Llyfr/Adroddiad/Trafodion Cynhadledd › Pennod › adolygiad gan gymheiriaid
- Cyhoeddwyd
Red squirrel population monitoring: Clocaenog forest
Shuttleworth, C. & Donoyou, T., 2012, IFCW108/001/002 gol.Allbwn ymchwil: Llyfr/Adroddiad › Adroddiad Comisiwn
- Cyhoeddwyd
Red squirrel conservation on Anglesey, North Wales. A report on the first five years of the Anglesey Red Squirrel Project
Shuttleworth, C., Bailey, M. & Knott, H., 2002Allbwn ymchwil: Llyfr/Adroddiad › Adroddiad Comisiwn
- Cyhoeddwyd
Red squirrel conservation and disease - Adenovirus in Wales
Shuttleworth, C., 2014, Yn: Natur Cymru. 52, t. 41-44Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl
- Cyhoeddwyd
Red squirrel bark-stripping of hornbeam in North Wales
Shuttleworth, C. & Gill, R., 2019, Yn: Quarterly Journal of Forestry. 113, 4, t. 275-277Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- Cyhoeddwyd
Red squirrel (Sciurus vulgaris) conservation plan for Anglesey & Gwynedd
Shuttleworth, C. & Halliwell, E. C., 2018, 50 t.Allbwn ymchwil: Llyfr/Adroddiad › Adroddiad Comisiwn
- Cyhoeddwyd
Red Squirrel Conservation in Clocaenog Forest: Review and Recommendations
Shuttleworth, C., 2021, 56 t.Allbwn ymchwil: Llyfr/Adroddiad › Adroddiad Comisiwn
- Cyhoeddwyd
Recalibrating Risk: Implications of squirrelpox virus for successful red squirrel translocations within mainland UK
Shuttleworth, C., Brady, D., Cross, P., Gardner, L., Greenwood, A., Jackson, N., McKinney, C., Robinson, N., Trotter, S., Valle, S., Wood, K. & Hayward, M., Chwef 2021, Yn: Conservation Science and Practice. 3, 2, e321.Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- Cyhoeddwyd
Re-introduction of the red squirrel into Newborough forest on the island of Anglesey, UK
Shuttleworth, C., Kenward, R. E. & Jackson, N., 2008, Global Re-Introduction Perspectives: Re-Introduction Case-Studies from Around the Globe. Soorae, P. S. (gol.). IUCN/SSC Re-introduction Specialist Group, t. 163-166Allbwn ymchwil: Pennod mewn Llyfr/Adroddiad/Trafodion Cynhadledd › Pennod
- Cyhoeddwyd
Rare woodland wildlife at risk because of 50-year-old tree felling rules
Shuttleworth, C., 2018, The Conversation.Allbwn ymchwil: Cyfraniad arall › Cyfraniad Arall
- Cyhoeddwyd
Ranging behaviour, density and social structure in grey squirrels
Lawton, C., Shuttleworth, C. & Kenward, R. E., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (gol.). European Squirrel Initiative, t. 133-152Allbwn ymchwil: Pennod mewn Llyfr/Adroddiad/Trafodion Cynhadledd › Pennod › adolygiad gan gymheiriaid