Dr Craig Shuttleworth
Honorary Visiting Research Fellow

Dolenni cyswllt
Contact info
Mobile 07966150847
Email c.shuttleworth@bangor.ac.uk
- Cyhoeddwyd
Preventing grey squirrel establishment on Anglesey
Shuttleworth, C., 2021, Invasive Alien Species Colonisation Prevention: Your guide to early detection and rapid response . Robinson, N. J. & Shuttleworth, C. M. (gol.). Royal Society of Wildlife Trusts, t. 55-69Allbwn ymchwil: Pennod mewn Llyfr/Adroddiad/Trafodion Cynhadledd › Pennod
- Cyhoeddwyd
Pine marten scat holds few clues for squirrel disease
Shuttleworth, C., Everest, D. J., Dastjerdi, A., Birks, J. & Hayward, M., 7 Rhag 2019, Yn: Veterinary Record. 185, t. 698Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl
- Cyhoeddwyd
Painting the Town Red: Gwynedd red squirrel conservation
Shuttleworth, C., 2015, Shared Experience of Red Squirrel Conservation Practice . Shuttleworth, C. M., Lurz, P. W. W. & Halliwell, E. C. (gol.). European Squirrel Initiative, t. 159-166Allbwn ymchwil: Pennod mewn Llyfr/Adroddiad/Trafodion Cynhadledd › Pennod
- Cyhoeddwyd
PCR detection of adenovirus in grey squirrels on Anglesey
Everest, D. J., Grierson, S. S., Stidworthy, M. & Shuttleworth, C., 31 Hyd 2009, Yn: Veterinary Record. 165, t. 482Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- Cyhoeddwyd
Opportunistic viral surveillance confirms the ongoing disease threat grey squirrels pose to sympatric red squirrel populations in the UK
Everest, D., Green, C., Dastjerdi, A., Davies, H., Cripps, R., McKinney, C., Podgornik, G., Stinson, M., O'Hare, S., Bonnie, S., Mill, A., Van der wall, Z., Robinson, N., Trotter, S. & Shuttleworth, C., 20 Mai 2023, Yn: Veterinary Record. 192, 10, e2834.Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- Cyhoeddwyd
Non-invasive genetic studies and virus epidemiology
Everest, D. J., Dastjerdi, A., Davies, H., Shuttleworth, C., Ogden, R., Butler, H. & Sapsford, B., 30 Maw 2022, Yn: Veterinary Record. 190, t. 204-205Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl
- Cyhoeddwyd
Monitoring escaped captive bred red squirrels in the UK
Shuttleworth, C., 2021, Saving the red squirrel: landscape recovery. Shuttleworth, C. M., Lurz, P. W. W. & Robinson, N. J. (gol.). Red Squirrel Survival Trust, t. 71-78Allbwn ymchwil: Pennod mewn Llyfr/Adroddiad/Trafodion Cynhadledd › Pennod
- Cyhoeddwyd
Mitochondrial DNA Analysis of Red Squirrels (Sciurus vulgaris) in North Wales
O’Meara, D., Shuttleworth, C., Clews-Roberts, R., Everest, D. & Synnott, R., 1 Maw 2025, 14 t. (Evidence Report Series; Rhif 791)Allbwn ymchwil: Llyfr/Adroddiad › Adroddiad Comisiwn
- Cyhoeddwyd
Mathematical models of grey squirrel invasion: a case study on Anglesey
Jones, H. E., White, A., Lurz, P. W. W., Boots, M. & Shuttleworth, C., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (gol.). European Squirrel Initiative, t. 235-255Allbwn ymchwil: Pennod mewn Llyfr/Adroddiad/Trafodion Cynhadledd › Pennod › adolygiad gan gymheiriaid
- Cyhoeddwyd
Mathematical models for invasive species management: grey squirrel control on Anglesey
Jones, H., White, A., Lurz, P. W. W. & Shuttleworth, C., 10 Medi 2017, Yn: Ecological Modelling. 180, t. 274-284Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- Cyhoeddwyd
Magical Mammals: Red squirrels in North West Wales
Shuttleworth, C., 2021, Saving the red squirrel: landscape recovery. Shuttleworth, C. M., Lurz, P. W. W. & Robinson, N. J. (gol.). Red Squirrel Survival Trust, t. 269-278Allbwn ymchwil: Pennod mewn Llyfr/Adroddiad/Trafodion Cynhadledd › Pennod
- Cyhoeddwyd
Leprosy in red squirrels in the UK
Schilling, A.-K., Del-Pozo, J., Lurz, P. W. W., Stevenson, K., Avanzi, C., Shuttleworth, C., Cole, S. T. & Meredith, A. L., 30 Maw 2019, Yn: Veterinary Record. 184, t. 416Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl
- Cyhoeddwyd
Landscape scale impacts of culling upon a European grey squirrel population: can trapping reduce population size and decrease the threat of squirrel pox virus infection for the native red squirrel?
Schuchert, P., Shuttleworth, C., McInnes, C. J., Everest, D. J. & Rushton, S. P., 1 Tach 2014, Yn: Biological Invasions. 16, t. 2381-2391Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- Cyhoeddwyd
Island Haven: Anglesey red squirrel conservation
Shuttleworth, C., 2015, Shared Experience of Red Squirrel Conservation Practice . Shuttleworth, C. M., Lurz, P. W. W. & Halliwell, E. C. (gol.). European Squirrel Initiative, t. 151-158Allbwn ymchwil: Pennod mewn Llyfr/Adroddiad/Trafodion Cynhadledd › Pennod
- Cyhoeddwyd
Invasive genetic rescue: dispersal following repeated culling reinforces the genetic diversity of an invasive mammal
Synnott, R., Shuttleworth, C., Everest, D., Stevenson-Holt, C., O'Reilly, C., McDevitt, A. & O.Meara, D., Hyd 2023, Yn: Biological Invasions. 25, t. 3323-3339Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- Cyhoeddwyd
Invasive alien species colonisation prevention: Your guide to early detection and rapid response
Robinson, N. & Shuttleworth, C., 2020, Newark: Royal Society of Wildlife Trusts. 204 t.Allbwn ymchwil: Llyfr/Adroddiad › Llyfr › adolygiad gan gymheiriaid
- Cyhoeddwyd
Introduced Canadian Eastern grey squirrels: squirrelpox virus surveillance and why nothing matters.
McInnes, C., Shuttleworth, C., Larsen, K. W., Everest, D. J., Bruemmer, C., Carroll, B., Romeo, C., Sainsbury, A. W., Crawshaw, G., Dubois, S., Gillis, L., Gilray, J. & Percival, A., 3 Awst 2020, Yn: Hystrix-Italian Journal of Mammalogy. 31, t. 95-98Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- Cyhoeddwyd
Interactions between the red squirrel (Sciurus vulgaris), great tit (Parus Major) and jackdaw (Corvus monedula) whilst using nest boxes.
Shuttleworth, C., 1 Hyd 2001, Yn: Journal of Zoology. 255, t. 269-272Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- Cyhoeddwyd
Inter-specific viral infections: Can the management of captive red squirrel collections help inform scientific research?
Shuttleworth, C., Everest, D. J., McInnes, C. J., Greenwood, A., Jackson, N., Rushton, S. P. & Kenward, R. E., 16 Meh 2014, Yn: Hystrix-Italian Journal of Mammalogy. 25, 1, t. 18-24Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- Cyhoeddwyd
Integrating red squirrel (Sciurus vulgaris) habitat requirements with the management of pathogenic tree disease in commercial forests in the UK
Shuttleworth, C., Lurz, P. W. W., Geddes, N. & Brown, J., 1 Medi 2012, Yn: Forest Ecology and Management. 279, t. 167-175Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- Cyhoeddwyd
Indirect effects of pine marten recovery result in benefits to native prey through suppression of an invasive species and a shared pathogen
Slade, A., White, A., Lurz, P. W. W., Shuttleworth, C., Tosh, D. & Twining, J., 1 Chwef 2023, Yn: Ecological Modelling. 476, 110216.Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- Cyhoeddwyd
Identifying incursion pathways, early detection responses and management actions to prevent grey squirrel range expansion: an island case study in Wales
Shuttleworth, C., Halliwell, E. C. & Robertson, P. A., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (gol.). European Squirrel Initiative, t. 475-494Allbwn ymchwil: Pennod mewn Llyfr/Adroddiad/Trafodion Cynhadledd › Pennod › adolygiad gan gymheiriaid
- Cyhoeddwyd
Gut bacteria may explain why grey squirrels outcompete reds – new research
Shuttleworth, C., 20 Chwef 2024, The Conversation.Allbwn ymchwil: Cyfraniad at gyhoeddiad arbenigol › Erthygl
- Cyhoeddwyd
Grey squirrels are bad for the British countryside – full stop
Shuttleworth, C., 2017, The Conversation.Allbwn ymchwil: Cyfraniad arall › Cyfraniad Arall
- Cyhoeddwyd
Grey squirrel nesting ecology and the use of nest sites in European population management
Shuttleworth, C., Selonen, V. & Koprowski, J. L., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (gol.). European Squirrel Initiative, t. 349-369Allbwn ymchwil: Pennod mewn Llyfr/Adroddiad/Trafodion Cynhadledd › Pennod › adolygiad gan gymheiriaid
- Cyhoeddwyd
Grey squirrel management in Europe - the future
Shuttleworth, C., Lurz, P. W. W. & Gurnell, J., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (gol.). European Squirrel InitiativeAllbwn ymchwil: Pennod mewn Llyfr/Adroddiad/Trafodion Cynhadledd › Pennod › adolygiad gan gymheiriaid
- Cyhoeddwyd
Grey squirrel control as a tactic in the conservation of the red squirrel: a positive outcome
Shuttleworth, C., 2001, Advances in Vertebrate Pest Management . Feare, C. J. & Cowan, D. P. (gol.). Filander Verlag, Cyfrol 3. t. 151-168Allbwn ymchwil: Pennod mewn Llyfr/Adroddiad/Trafodion Cynhadledd › Pennod › adolygiad gan gymheiriaid
- Cyhoeddwyd
Genetic management of the red squirrel, Sciurus vulgaris: a practical approach to regional conservation
Ogden, R., Shuttleworth, C., McEwing, R. & Cesarini, S., 26 Hyd 2005, Yn: Conservation Genetics. 6, t. 511-525Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- Cyhoeddwyd
Genetic evidence of the effectiveness of grey squirrel control operations: lessons from the Isle of Anglesey
Signorile, L. & Shuttleworth, C., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (gol.). European Squirrel Initiative, t. 439-452Allbwn ymchwil: Pennod mewn Llyfr/Adroddiad/Trafodion Cynhadledd › Pennod › adolygiad gan gymheiriaid
- Cyhoeddwyd
Future red squirrel conservation in Clocaenog forest
Shuttleworth, C. & Hayward, M., 2015, P21018-0026 gol.Allbwn ymchwil: Llyfr/Adroddiad › Adroddiad Comisiwn
- Cyhoeddwyd
First report of dermatophilosis in wild European red squirrels (Sciurus vulgaris)
Holmes, P., Everest, D. J., Spiro, S., Wessels, M. & Shuttleworth, C., 30 Medi 2019, Yn: Veterinary Record Case Reports. 7, 3, t. 3Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- Cyhoeddwyd
Experiences of grey squirrel management in an upland conifer forest
Halliwell, E. C., Shuttleworth, C., Cartmel, S., Lloyd, I. & Jenkins, R., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (gol.). European Squirrel Initiative, t. 453-474Allbwn ymchwil: Pennod mewn Llyfr/Adroddiad/Trafodion Cynhadledd › Pennod › adolygiad gan gymheiriaid
- Cyhoeddwyd
Evolving grey squirrel management techniques in Europe
Shuttleworth, C., Robinson, N. J., Halliwell, E. C., Clews-Roberts, R., Peek, H., Podgornik, G., Stinson, M., Rice, S., Finlay, C., McKinney, C., Everest, D. J. & Larsen, K. W., 21 Hyd 2020, Yn: Management of Biological Invasions. 11, 4, t. 747-761Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- Cyhoeddwyd
Ethical dilemmas when using citizen science for early detection of invasive tree pests and diseases
Pocock, M. J. O., Marzano, M., Bullas-Appleton, E., Dyke, A., de Groot, M., Shuttleworth, C. & White, R., 1 Tach 2020, Yn: Management of Biological Invasions. 11, t. 720-732Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- Cyhoeddwyd
Envisioning the future with 'compassionate conservation': An ominous projection for native wildlife and biodiversity
Callen, A., Hayward, M. W., Klop-Toker, K., Allen, B. L., Ballard, G., Beranek, C. T., Broekhuis, F., Bugir, C. K., Clarke, R. H., Clulow, J., Clulow, S., Daltry, J. C., Davies-Mostert, H. T., Di Blanco, Y. E., Dixon, V., Fleming, P. J. S., Howell, L. G., Kerley, G. I. H., Legge, S. M., Lenga, D. J., Major, T., Montgomery, R. A., Moseby, K., Meyer, N., Parker, D. M., Periquet, S., Read, J., Scanlon, R. J., Shuttleworth, C., Tamessar, C. T., Taylor, W. A., Tuft, K., Upton, R. M. O., Valenzuela, M., Witt, R. R. & Wuster, W., Ion 2020, Yn: Biological Conservation. 241, 108365.Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- Cyhoeddwyd
Ecosystem impacts of an alien invader in Europe, the grey squirrel Sciurus carolinensis
Gurnell, J., Lurz, P. W. W. & Shuttleworth, C., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (gol.). European Squirrel Initiative, t. 307-329Allbwn ymchwil: Pennod mewn Llyfr/Adroddiad/Trafodion Cynhadledd › Pennod › adolygiad gan gymheiriaid
- Cyhoeddwyd
Do differing levels of boldness influence the success of translocation? A pilot study on red squirrels (Sciurus vulgaris)
Bamber, J. A., Shuttleworth, C. & Hayward, M., 26 Medi 2020, Yn: Animals. 10, 1748.Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- Cyhoeddwyd
Disease, invasions and conservation: no evidence of squirrelpox virus in grey squirrels introduced to Italy
Romeo, C., McInnes, C., Dale, T., Shuttleworth, C., Bertolino, S., Wauters, L. A. & Ferrari, N., 1 Chwef 2019, Yn: Animal Conservation. 22, 1, t. 14-23Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- Cyhoeddwyd
Disease monitoring and surveillance: case studies in the applied conservation of fragmented red squirrel (Sciurus vulgaris) populations in England and Wales
Everest, D. J., Floyd, T., Holmes, P., Man, C., Dunnett, C., Locke, R., Savage, L., Sutcliffe, S., Sapsford, B., Shuttleworth, C. & Duff, P., 1 Rhag 2021, Yn: Mammalian Biology. 101, t. 1079-1088Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- Cyhoeddwyd
Developing red squirrel re-introduction techniques for use during regional grey squirrel eradication programmes in Europe
Shuttleworth, C., Jackson, N. & Kenward, R. E., 2015, 2015 Global Reintroduction Perspectives. Soorae, P. (gol.). IUCN/SSC Re-introduction Specialist Group, t. 67-74Allbwn ymchwil: Pennod mewn Llyfr/Adroddiad/Trafodion Cynhadledd › Pennod
- Cyhoeddwyd
Developing integrated and applied red squirrel conservation programmes: What lessons can Europe learn from a regional grey squirrel eradication programme in North Wales?
Shuttleworth, C., Schuchert, P., Everest, D. J., McInnes, C. J., Rushton, S. P., Jackson, N. & Kenward, R. E., 2015, Red squirrels: Ecology, Conservation & Management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Hayward, M. W. (gol.). European Squirrel Initiative, t. 233-250Allbwn ymchwil: Pennod mewn Llyfr/Adroddiad/Trafodion Cynhadledd › Pennod › adolygiad gan gymheiriaid
- Cyhoeddwyd
Developing a method for assessing the relative abundance of red squirrels in low-density populations
Shannon, G., Barton, O., Valle, S. & Shuttleworth, C., 1 Tach 2024, 34 t.Allbwn ymchwil: Llyfr/Adroddiad › Adroddiad Comisiwn
- Cyhoeddwyd
Detecting viral infection in red squirrels
Shuttleworth, C., Everest, D. J., Halliwell, E. C., Hulme, B., Wilberforce, L. & Clews-Roberts, R., 30 Ebr 2019, Yn: Veterinary Record. 184, t. 507Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl
- Cyhoeddwyd
Deconstructing compassionate conservation
Hayward, M. W., Callen, A., Allen, B. L., Ballard, G., Broekhuis, F., Bugir, C., Clarke, R. H., Clulow, J., Clulow, S., Daltry, J. C., Davies-Mostert, H. T., Fleming, P. J. S., Griffin, A. S., Howell, L. G., Kerley, G. I. H., Klop-Toker, K., Legge, S., Major, T., Meyer, N., Montgomery, R. A., Moseby, K., Parker, D. M., Périquet, S., Read, J., Scanlon, R., Seeto, R., Shuttleworth, C., Somers, M. J., Tamessar, C. T., Tuft, K., Upton, R., Valenzuela-Molina, M., Wayne, A., Witt, R. R. & Wuster, W., Awst 2019, Yn: Conservation Biology. 33, 4, t. 760-768Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- Cyhoeddwyd
Controlling grey squirrels in forests and woodlands in the UK
Gill, R., Ferryman, M., Shuttleworth, C., Lurz, P. W. W., Mill, A., Robertson, P. A. & Dutton, C., 2019, Forest Research. 16 t.Allbwn ymchwil: Llyfr/Adroddiad › Llyfr › adolygiad gan gymheiriaid
- Cyhoeddwyd
Confirmation of squirrelpox in Welsh red squirrel
Everest, D. J., Floyd, T., Donnnacie, B., Irvine, R. M., Holmes, J. P. & Shuttleworth, C., 30 Tach 2017, Yn: Veterinary Record. 181, t. 514-515Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl
- Cyhoeddwyd
Challenges and solutions in early detection, rapid response and communication about potential invasive alien species in forests
de Groot, M., Bullas-Appleton, E., Csóka, G., Csiszár, A., Faccoli, A., Gervasini, A., Kirichenko, N., Shuttleworth, C., Kus Veenvliet, J., O'Hanlon, R., Korda, M., Marinsek, A., Robinson, N., Sweeney, J., Tricarico, E., Verbrugge, L., Williams, D. & Zidar, S., 14 Awst 2020, Yn: Management of Biological Invasions. 11, t. 637-660Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- Cyhoeddwyd
Capturing red squirrels (Sciurus vulgaris) on camera: A cost-effective approach for monitoring relative abundance and habitat preference
Shannon, G., Valle, S. & Shuttleworth, C., 3 Hyd 2023, Yn: Ecology and Evolution. 13, 10, e10536.Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
- Cyhoeddwyd
By trying to be everything, rewilding risks meaning nothing
Jachowski, J. D., Bombaci, S., Caravaggi, A., Chalmers, A., Clulow, S., Griffin, A., Hayward, M., Heurich, M., Linnell, J., Marnewick, K., Moehrenschlager, A., Montgomery, A., Shuttleworth, C., Somers, M. & Weise, F., Tach 2019, Yn: Science. 364, t. 6438Allbwn ymchwil: Cyfraniad at gyfnodolyn › Llythyr
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British red squirrels remain the only known wild rodent host for leprosy bacilli
Schilling, A., Avanzi, C., Ulrich, R. G., Busso, P., Pisanu, B., Ferrari, N., Romeo, C., Mazzamuto, M. V., McLuckie, J., Shuttleworth, C., Del-Pozo, J., Lurz, P. W. W., Escalante-Fuentes, W. G., Ocampo-Candiani, J., Vera-Cabrera, L., Stevenson, K., Chapuis, J.-L., Meredith, A. L. & Cole, S. T., 1 Chwef 2019, Yn: Frontiers in Veterinary Science. 6, t. 8Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid