Dr Craig Shuttleworth
Honorary Visiting Research Fellow

Affiliations
Links
- https://twitter.com/Welshsquirrels
Personal account - https://twitter.com/RedSquirrelsinW
Welsh red squirrel conservation - https://www.facebook.com/groups/143610382316300
Welsh red squirrel conservation - https://pinemarten.net/
The Gwynedd pine marten project - https://www.linkedin.com/in/craig-shuttleworth-6062a1253/
Personal page
Contact info
Mobile 07966150847
Email c.shuttleworth@bangor.ac.uk
- Published
Landscape scale impacts of culling upon a European grey squirrel population: can trapping reduce population size and decrease the threat of squirrel pox virus infection for the native red squirrel?
Schuchert, P., Shuttleworth, C., McInnes, C. J., Everest, D. J. & Rushton, S. P., 1 Nov 2014, In: Biological Invasions. 16, p. 2381-2391Research output: Contribution to journal › Article › peer-review
- Published
Capturing red squirrels (Sciurus vulgaris) on camera: A cost-effective approach for monitoring relative abundance and habitat preference
Shannon, G., Valle, S. & Shuttleworth, C., 3 Oct 2023, In: Ecology and Evolution. 13, 10, e10536.Research output: Contribution to journal › Article › peer-review
- Published
Developing a method for assessing the relative abundance of red squirrels in low-density populations
Shannon, G., Barton, O., Valle, S. & Shuttleworth, C., 1 Nov 2024, 34 p.Research output: Book/Report › Commissioned report
- Published
The grey squirrel: ecology & management of an Invasive Species in Europe
Shuttleworth, C., Lurz, P. W. W. & Gurnell, J., 2016, European Squirrel Initiative. 505 p.Research output: Book/Report › Book › peer-review
- Published
The foraging behaviour and diet of red squirrels (Sciurus vulgaris) receiving supplemental feeding
Shuttleworth, C., 1 Sept 2000, In: Wildlife Biology. 6, p. 149-156Research output: Contribution to journal › Article › peer-review
- Published
The management of coastal sand dune woodland for red squirrels (Sciurus vulgaris L.)
Shuttleworth, C. & Gurnell, J., 2001, Coastal Dune Management: Shared Experience of European Conservation Practice. Houston, J., Edmondson, A. S. E. & Rooney, P. J. (eds.). Liverpool University Press, p. 117-127Research output: Chapter in Book/Report/Conference proceeding › Chapter
- Published
Traffic related mortality in a red squirrel (Sciurus vulgaris) population receiving supplemental feeding
Shuttleworth, C., 1 Jun 2001, In: Urban Ecosystems. 2, p. 109-118Research output: Contribution to journal › Article › peer-review
- Published
Preventing grey squirrel establishment on Anglesey
Shuttleworth, C., 2021, Invasive Alien Species Colonisation Prevention: Your guide to early detection and rapid response . Robinson, N. J. & Shuttleworth, C. M. (eds.). Royal Society of Wildlife Trusts, p. 55-69Research output: Chapter in Book/Report/Conference proceeding › Chapter
- Published
Painting the Town Red: Gwynedd red squirrel conservation
Shuttleworth, C., 2015, Shared Experience of Red Squirrel Conservation Practice . Shuttleworth, C. M., Lurz, P. W. W. & Halliwell, E. C. (eds.). European Squirrel Initiative, p. 159-166Research output: Chapter in Book/Report/Conference proceeding › Chapter
- Published
Magical Mammals: Red squirrels in North West Wales
Shuttleworth, C., 2021, Saving the red squirrel: landscape recovery. Shuttleworth, C. M., Lurz, P. W. W. & Robinson, N. J. (eds.). Red Squirrel Survival Trust, p. 269-278Research output: Chapter in Book/Report/Conference proceeding › Chapter
- Published
Adenovirus remains widespread in squirrels on Anglesey
Shuttleworth, C., Everest, D. & McQueen, A., 16 Dec 2023, In: Veterinary Record. 193, 12, p. 490-491 2 p.Research output: Contribution to journal › Letter
- Published
Red squirrel conservation and disease - Adenovirus in Wales
Shuttleworth, C., 2014, In: Natur Cymru. 52, p. 41-44Research output: Contribution to journal › Article
- Published
Identifying incursion pathways, early detection responses and management actions to prevent grey squirrel range expansion: an island case study in Wales
Shuttleworth, C., Halliwell, E. C. & Robertson, P. A., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (eds.). European Squirrel Initiative, p. 475-494Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review
- Published
Red squirrel population monitoring: Clocaenog forest
Shuttleworth, C. & Donoyou, T., 2012, IFCW108/001/002 ed.Research output: Book/Report › Commissioned report
- Published
Shared experience of red squirrel conservation practice
Shuttleworth, C., Lurz, P. W. W. & Halliwell, E. C., 2015, European Squirrel Initiative. 220 p.Research output: Book/Report › Book
- Published
Island Haven: Anglesey red squirrel conservation
Shuttleworth, C., 2015, Shared Experience of Red Squirrel Conservation Practice . Shuttleworth, C. M., Lurz, P. W. W. & Halliwell, E. C. (eds.). European Squirrel Initiative, p. 151-158Research output: Chapter in Book/Report/Conference proceeding › Chapter
- Published
Gut bacteria may explain why grey squirrels outcompete reds – new research
Shuttleworth, C., 20 Feb 2024, The Conversation.Research output: Contribution to specialist publication › Article
- Published
Turning the grey tide: progress in red squirrel recovery
Shuttleworth, C., 1 Jun 2010, In: Ecos. 31, p. 27-35Research output: Contribution to journal › Article
- Published
The role of spruce in red squirrel (Sciurus vulgaris) conservation in the UK
Shuttleworth, C. & Lurz, P. W. W., 2014, Forest Ecosystems: Biodiversity, Management and Conservation. Roberts, N. C. (ed.). Nova Science Publishers, p. 63-78Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review
- Published
Saving the red squirrel: landscape scale recovery
Shuttleworth, C., Lurz, P. W. W. & Robinson, N., 2021, London: Red Squirrel Survival Trust. 306 p.Research output: Book/Report › Book › peer-review
- Published
Red squirrel conservation on Anglesey, North Wales. A report on the first five years of the Anglesey Red Squirrel Project
Shuttleworth, C., Bailey, M. & Knott, H., 2002Research output: Book/Report › Commissioned report
- Published
Reducing negative impacts on non-target species during grey squirrel (Sciurus carolinensis) control activities
Shuttleworth, C., Mill, A. C., Van der Wall, Z., Bertolino, S. & Everest, D. J., 1 Jan 2018, In: Quarterly Journal of Forestry. 112, 1, p. 21-29Research output: Contribution to journal › Article › peer-review
- Published
The effect of supplemental feeding on the diet, population density and reproduction of red squirrels
Shuttleworth, C., 1997, The Conservation of Red Squirrels (Sciurus vulgaris L.). Gurnell, J. & Lurz, P. W. W. (eds.). Peoples Trust for Endangered Species, p. 13-24Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review
- Published
Releasing grey squirrels into the wild
Shuttleworth, C., Bertolino, S., Gill, R., Gurnell, J., Hayward, M., Kenward, R. E., Lawton, C., Lurz, P. W. W., McInnes, C. J., Mill, A., Trotter, S. & Wauters, L. A., 31 Mar 2019, In: Veterinary Record. 184, p. 389-390Research output: Contribution to journal › Article
- Published
Interactions between the red squirrel (Sciurus vulgaris), great tit (Parus Major) and jackdaw (Corvus monedula) whilst using nest boxes.
Shuttleworth, C., 1 Oct 2001, In: Journal of Zoology. 255, p. 269-272Research output: Contribution to journal › Article › peer-review
- Published
Inter-specific viral infections: Can the management of captive red squirrel collections help inform scientific research?
Shuttleworth, C., Everest, D. J., McInnes, C. J., Greenwood, A., Jackson, N., Rushton, S. P. & Kenward, R. E., 16 Jun 2014, In: Hystrix-Italian Journal of Mammalogy. 25, 1, p. 18-24Research output: Contribution to journal › Article › peer-review
- Published
Are nest boxes a useful tool in regional red squirrel conservation programs?
Shuttleworth, C. & Schuchert , P., 15 Dec 2014, In: Hystrix-Italian Journal of Mammalogy. 25, p. 91-94Research output: Contribution to journal › Article › peer-review
- Published
Re-introduction of the red squirrel into Newborough forest on the island of Anglesey, UK
Shuttleworth, C., Kenward, R. E. & Jackson, N., 2008, Global Re-Introduction Perspectives: Re-Introduction Case-Studies from Around the Globe. Soorae, P. S. (ed.). IUCN/SSC Re-introduction Specialist Group, p. 163-166Research output: Chapter in Book/Report/Conference proceeding › Chapter
- Published
Evolving grey squirrel management techniques in Europe
Shuttleworth, C., Robinson, N. J., Halliwell, E. C., Clews-Roberts, R., Peek, H., Podgornik, G., Stinson, M., Rice, S., Finlay, C., McKinney, C., Everest, D. J. & Larsen, K. W., 21 Oct 2020, In: Management of Biological Invasions. 11, 4, p. 747-761Research output: Contribution to journal › Article › peer-review
- Published
The reintroduction of the red squirrel Sciurus vulgaris to Newborough forest, North Wales: A five year project review
Shuttleworth, C., Kenward, R. E. & Jackson, N., 2009Research output: Book/Report › Commissioned report
- Published
The impact of woodland management upon the red squirrel population in Ainsdale National Nature Reserve
Shuttleworth, C., 1997Research output: Book/Report › Commissioned report
- Published
Red squirrels in my garden: guidance and tips to help conserve local populations
Shuttleworth, C. & Halliwell, E. C., 2018, European Squirrel Initiative. 185 p.Research output: Book/Report › Book › peer-review
- Published
Against a grey tide: What future for the red squirrel?
Shuttleworth, C., 2005, Journal of Practical Ecology & Conservation (Special Series No 4): Living with the Aliens? - Ecology, History, Culture and Management if exotic plants and animals: issues for nature conservation . Rotherham, L. D. (ed.). Vol. 4. p. 87-88Research output: Chapter in Book/Report/Conference proceeding › Chapter
- Published
Squirrelpox outbreak detected in north Wales – without a vaccine, the disease will keep decimating red squirrels
Shuttleworth, C., 21 Dec 2022, The Conversation.Research output: Other contribution
- Published
Grey squirrel management in Europe - the future
Shuttleworth, C., Lurz, P. W. W. & Gurnell, J., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (eds.). European Squirrel InitiativeResearch output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review
- Published
Monitoring escaped captive bred red squirrels in the UK
Shuttleworth, C., 2021, Saving the red squirrel: landscape recovery. Shuttleworth, C. M., Lurz, P. W. W. & Robinson, N. J. (eds.). Red Squirrel Survival Trust, p. 71-78Research output: Chapter in Book/Report/Conference proceeding › Chapter
- Published
Red squirrels (Sciurus vulgaris) on the Sefton Coast: The conservation value of nest boxes
Shuttleworth, C., 1995, 2nd NPI Red Alert UK Forum for Red Squirrel Conservation. Hughes, D. G. & Tew, T. (eds.). The Federation of Zoological Gardens of Great Britain and Ireland, p. 25-32Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review
- Published
Red squirrel bark-stripping of hornbeam in North Wales
Shuttleworth, C. & Gill, R., 2019, In: Quarterly Journal of Forestry. 113, 4, p. 275-277Research output: Contribution to journal › Article › peer-review
- Published
Red squirrels: ecology, conservation & management in Europe
Shuttleworth, C., Hayward, M. & Lurz, P. W. W., 2015, European Squirrel Initiative. 330 p.Research output: Book/Report › Book › peer-review
- Published
Pine marten scat holds few clues for squirrel disease
Shuttleworth, C., Everest, D. J., Dastjerdi, A., Birks, J. & Hayward, M., 7 Dec 2019, In: Veterinary Record. 185, p. 698Research output: Contribution to journal › Article
- Published
Recalibrating Risk: Implications of squirrelpox virus for successful red squirrel translocations within mainland UK
Shuttleworth, C., Brady, D., Cross, P., Gardner, L., Greenwood, A., Jackson, N., McKinney, C., Robinson, N., Trotter, S., Valle, S., Wood, K. & Hayward, M., Feb 2021, In: Conservation Science and Practice. 3, 2, e321.Research output: Contribution to journal › Article › peer-review
- Published
Integrating red squirrel (Sciurus vulgaris) habitat requirements with the management of pathogenic tree disease in commercial forests in the UK
Shuttleworth, C., Lurz, P. W. W., Geddes, N. & Brown, J., 1 Sept 2012, In: Forest Ecology and Management. 279, p. 167-175Research output: Contribution to journal › Article › peer-review
- Published
Wales should follow Scotland’s lead in protecting the red squirrel’s habitat
Shuttleworth, C., 2018, Nation Cymru .Research output: Other contribution
- Published
Rare woodland wildlife at risk because of 50-year-old tree felling rules
Shuttleworth, C., 2018, The Conversation.Research output: Other contribution
- Published
Developing red squirrel re-introduction techniques for use during regional grey squirrel eradication programmes in Europe
Shuttleworth, C., Jackson, N. & Kenward, R. E., 2015, 2015 Global Reintroduction Perspectives. Soorae, P. (ed.). IUCN/SSC Re-introduction Specialist Group, p. 67-74Research output: Chapter in Book/Report/Conference proceeding › Chapter
- Published
An Opportunistic Assessment of the Impact of Squirrelpox Disease Outbreaks upon a Red Squirrel Population Sympatric with Grey Squirrels in Wales
Shuttleworth, C., Everest, D., Holmes, P., Bell, S. & Cripps, R., 1 Jan 2022, In: Animals. 12, 1Research output: Contribution to journal › Article › peer-review
- Published
The use of nest boxes by the red squirrel (Sciurus vulgaris) in conifer plantations
Shuttleworth, C., 2001, In: Quarterly Journal of Forestry. 95, p. 225-228Research output: Contribution to journal › Article › peer-review
- Published
Why the pine marten is not every red squirrel’s best friend
Shuttleworth, C. & Hayward, M., 2019, The Conversation.Research output: Other contribution
- Published
Grey squirrel nesting ecology and the use of nest sites in European population management
Shuttleworth, C., Selonen, V. & Koprowski, J. L., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (eds.). European Squirrel Initiative, p. 349-369Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review
- Published
The use of nest boxes by the red squirrel Sciurus vulgaris in a coniferous habitat
Shuttleworth, C., 1 Mar 1999, In: Mammal Review. 29, p. 61-66Research output: Contribution to journal › Article › peer-review