Dr Craig Shuttleworth
Honorary Visiting Research Fellow

Affiliations
Links
- https://twitter.com/Welshsquirrels
Personal account - https://twitter.com/RedSquirrelsinW
Welsh red squirrel conservation - https://www.facebook.com/groups/143610382316300
Welsh red squirrel conservation - https://pinemarten.net/
The Gwynedd pine marten project - https://www.linkedin.com/in/craig-shuttleworth-6062a1253/
Personal page
Contact info
Mobile 07966150847
Email c.shuttleworth@bangor.ac.uk
- 2016
- Published
Grey squirrel nesting ecology and the use of nest sites in European population management
Shuttleworth, C., Selonen, V. & Koprowski, J. L., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (eds.). European Squirrel Initiative, p. 349-369Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review
- Published
Identifying incursion pathways, early detection responses and management actions to prevent grey squirrel range expansion: an island case study in Wales
Shuttleworth, C., Halliwell, E. C. & Robertson, P. A., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (eds.). European Squirrel Initiative, p. 475-494Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review
- Published
Mathematical models of grey squirrel invasion: a case study on Anglesey
Jones, H. E., White, A., Lurz, P. W. W., Boots, M. & Shuttleworth, C., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (eds.). European Squirrel Initiative, p. 235-255Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review
- Published
Ranging behaviour, density and social structure in grey squirrels
Lawton, C., Shuttleworth, C. & Kenward, R. E., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (eds.). European Squirrel Initiative, p. 133-152Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review
- Published
The grey squirrel: ecology & management of an Invasive Species in Europe
Shuttleworth, C., Lurz, P. W. W. & Gurnell, J., 2016, European Squirrel Initiative. 505 p.Research output: Book/Report › Book › peer-review
- Published
The management of grey squirrel populations in Europe: evolving best practice
Bertolino, S., Lurz, P. W. W., Shuttleworth, C., Martinoli, A. & Wauters, L. A., 2016, The grey squirrel: ecology and management in Europe . Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (eds.). European Squirrel Initiative, p. 495-516Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review
- Published
The potential role of ectoparasites in the epidemiology of squirrelpox virus: a possible novel means of intervention to reduce the impact of the disease?
Cowan, D. P., Mill, A. C., Everest, D. J., McInnes, C. J., Rushton, S. P., Shirley, M. D. F., Start, C. & Shuttleworth, C., 2016, The grey squirrel: ecology and management in Europe . Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (eds.). European Squirrel Initiative, p. 255-289Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review
- Published
A preliminary comparison of magnum 116 body-grip traps with a live capture trap design during the control of grey squirrels (Sciurus carolinensis)
Shuttleworth, C., Mill, A. & Van der Waal, Z., Dec 2016, In: International Journal of Pest Management. 63, 4, p. 316-321Research output: Contribution to journal › Article › peer-review
- 2017
- Published
Grey squirrels are bad for the British countryside – full stop
Shuttleworth, C., 2017, The Conversation.Research output: Other contribution
- Published
The large-scale removal of mammalian invasive alien species in Northern Europe
Robertson, P. A., Adriaens, T., Lambin, X., Mill, A., Roy, S., Shuttleworth, C. & Sutton-Croft, M., 1 Feb 2017, In: Pest Management Science. 73, p. 273-279Research output: Contribution to journal › Article › peer-review
- Published
Virus presence within Welsh red squirrels
Everest, D. J., Holmes, P. & Shuttleworth, C., 30 May 2017, In: The Veterinary record. 180, p. 520-521Research output: Contribution to journal › Article
- Published
Mathematical models for invasive species management: grey squirrel control on Anglesey
Jones, H., White, A., Lurz, P. W. W. & Shuttleworth, C., 10 Sept 2017, In: Ecological Modelling. 180, p. 274-284Research output: Contribution to journal › Article › peer-review
- Published
Confirmation of squirrelpox in Welsh red squirrel
Everest, D. J., Floyd, T., Donnnacie, B., Irvine, R. M., Holmes, J. P. & Shuttleworth, C., 30 Nov 2017, In: Veterinary Record. 181, p. 514-515Research output: Contribution to journal › Article
- 2018
- Published
Rare woodland wildlife at risk because of 50-year-old tree felling rules
Shuttleworth, C., 2018, The Conversation.Research output: Other contribution
- Published
Red squirrel (Sciurus vulgaris) conservation plan for Anglesey & Gwynedd
Shuttleworth, C. & Halliwell, E. C., 2018, 50 p.Research output: Book/Report › Commissioned report
- Published
Red squirrels in my garden: guidance and tips to help conserve local populations
Shuttleworth, C. & Halliwell, E. C., 2018, European Squirrel Initiative. 185 p.Research output: Book/Report › Book › peer-review
- Published
Wales should follow Scotland’s lead in protecting the red squirrel’s habitat
Shuttleworth, C., 2018, Nation Cymru .Research output: Other contribution
- Published
Reducing negative impacts on non-target species during grey squirrel (Sciurus carolinensis) control activities
Shuttleworth, C., Mill, A. C., Van der Wall, Z., Bertolino, S. & Everest, D. J., 1 Jan 2018, In: Quarterly Journal of Forestry. 112, 1, p. 21-29Research output: Contribution to journal › Article › peer-review
- Published
The implications of significant adenovirus infection in UK captive red squirrel (Sciurus vulgaris) collections: How histological screening can aid applied conservation management
Everest, D. J., Shuttleworth, C., Grierson, S. S., Dastjerdi, A., M.F., S., Duff, J. P., Higgins, R. J., Mill, A. C. & Chantrey, J., 1 Jan 2018, In: Mammalian Biology. 88, p. 123-129Research output: Contribution to journal › Article › peer-review
- 2019
- Published
Controlling grey squirrels in forests and woodlands in the UK
Gill, R., Ferryman, M., Shuttleworth, C., Lurz, P. W. W., Mill, A., Robertson, P. A. & Dutton, C., 2019, Forest Research. 16 p.Research output: Book/Report › Book › peer-review
- Published
Red squirrel bark-stripping of hornbeam in North Wales
Shuttleworth, C. & Gill, R., 2019, In: Quarterly Journal of Forestry. 113, 4, p. 275-277Research output: Contribution to journal › Article › peer-review
- Published
Why the pine marten is not every red squirrel’s best friend
Shuttleworth, C. & Hayward, M., 2019, The Conversation.Research output: Other contribution
- Published
British red squirrels remain the only known wild rodent host for leprosy bacilli
Schilling, A., Avanzi, C., Ulrich, R. G., Busso, P., Pisanu, B., Ferrari, N., Romeo, C., Mazzamuto, M. V., McLuckie, J., Shuttleworth, C., Del-Pozo, J., Lurz, P. W. W., Escalante-Fuentes, W. G., Ocampo-Candiani, J., Vera-Cabrera, L., Stevenson, K., Chapuis, J.-L., Meredith, A. L. & Cole, S. T., 1 Feb 2019, In: Frontiers in Veterinary Science. 6, p. 8Research output: Contribution to journal › Article › peer-review
- Published
Disease, invasions and conservation: no evidence of squirrelpox virus in grey squirrels introduced to Italy
Romeo, C., McInnes, C., Dale, T., Shuttleworth, C., Bertolino, S., Wauters, L. A. & Ferrari, N., 1 Feb 2019, In: Animal Conservation. 22, 1, p. 14-23Research output: Contribution to journal › Article › peer-review
- Published
Leprosy in red squirrels in the UK
Schilling, A.-K., Del-Pozo, J., Lurz, P. W. W., Stevenson, K., Avanzi, C., Shuttleworth, C., Cole, S. T. & Meredith, A. L., 30 Mar 2019, In: Veterinary Record. 184, p. 416Research output: Contribution to journal › Article
- Published
Releasing grey squirrels into the wild
Shuttleworth, C., Bertolino, S., Gill, R., Gurnell, J., Hayward, M., Kenward, R. E., Lawton, C., Lurz, P. W. W., McInnes, C. J., Mill, A., Trotter, S. & Wauters, L. A., 31 Mar 2019, In: Veterinary Record. 184, p. 389-390Research output: Contribution to journal › Article
- Published
Detecting viral infection in red squirrels
Shuttleworth, C., Everest, D. J., Halliwell, E. C., Hulme, B., Wilberforce, L. & Clews-Roberts, R., 30 Apr 2019, In: Veterinary Record. 184, p. 507Research output: Contribution to journal › Article
- Published
Reintroducing rewilding to restoration – Rejecting the search for novelty
Hayward, M. W., Scanlon, R. J., Callen, A., Howell, L. G., Klop-Toker, K. L., Di Blanco, Y., Balkenhol, N., Bugir, C. K., Campbell, L., Caravaggi, A., Chalmers, A. C., Clulow, J., Clulow, S., Cross, P., Gould, J. A., Griffin, A. S., Heurich, M., Howe, B. K., Jachowski, D. S., Jhala, Y. V., Krishnamurthy, R., Kowalczyk, R., Lenga, D. J., Linnell, J. D. C., Marnewick, K. A., Moehrenschlager, A., Montgomery, R. A., Osipova, L., Peneaux, C., Rodger, J. C., Sales, L. P., Seeto, R. G. Y., Shuttleworth, C. M., Somers, M. J., Tamessar, C. T., Upton, R. M. O. & Weise, F. J., 1 May 2019, In: Biological Conservation. 233, p. 255-259 5 p.Research output: Contribution to journal › Article › peer-review
- Published
Rewilding needs to clarify the role of management of invasive species
Hayward, M., Jachowski, D. S., Shuttleworth, C., Linnell, J. D. C., Allen, B. L., Griffen, A. S., Montgomery, R. A., Caravaggi, A., Weise, F. J., Moehrenschlager, A., Somers, M. J., Clulow, S., Heurich, M. & Marnewick, K., 8 May 2019, In: Science. 364, p. 6438Research output: Contribution to journal › Letter
- Published
Assessing a potential non-invasive method for viral diagnostic purposes in European squirrels
Everest, D. J., Tolhurst-Cherriman, D. A. R., Davies, H., Dastjerdi, A., Ashton, A., Blackett, T., Meredith, A. L., Milne, E., Mill, A. & Shuttleworth, C., 4 Jun 2019, In: Hystrix-Italian Journal of Mammalogy. 30, p. 44-50Research output: Contribution to journal › Article › peer-review
- Published
Deconstructing compassionate conservation
Hayward, M. W., Callen, A., Allen, B. L., Ballard, G., Broekhuis, F., Bugir, C., Clarke, R. H., Clulow, J., Clulow, S., Daltry, J. C., Davies-Mostert, H. T., Fleming, P. J. S., Griffin, A. S., Howell, L. G., Kerley, G. I. H., Klop-Toker, K., Legge, S., Major, T., Meyer, N., Montgomery, R. A., Moseby, K., Parker, D. M., Périquet, S., Read, J., Scanlon, R., Seeto, R., Shuttleworth, C., Somers, M. J., Tamessar, C. T., Tuft, K., Upton, R., Valenzuela-Molina, M., Wayne, A., Witt, R. R. & Wuster, W., Aug 2019, In: Conservation Biology. 33, 4, p. 760-768Research output: Contribution to journal › Article › peer-review
- Published
The search for novelty continues for rewilding
Hayward, M., Jachowski, J. D., Bugir, C. J., Clulow, J., Krishnamurthy, R., Griffin, A. S., Chalmers, A. C., Linnell, J. D. C., Montgomery, R. A., Somers, M. J., Kowalczyk, R., Heurich, M., Caravaggi, A., Marnewick, K. A., Di Blanco, Y., Shuttleworth, C., Callen, C., Weise, F., Scanlon, R., Moehrenschlager, A., Howell, L. & Upton, R. M. O., 1 Aug 2019, In: Biological Conservation. 236, p. 584-585Research output: Contribution to journal › Article › peer-review
- Published
First report of dermatophilosis in wild European red squirrels (Sciurus vulgaris)
Holmes, P., Everest, D. J., Spiro, S., Wessels, M. & Shuttleworth, C., 30 Sept 2019, In: Veterinary Record Case Reports. 7, 3, p. 3Research output: Contribution to journal › Article › peer-review
- Published
By trying to be everything, rewilding risks meaning nothing
Jachowski, J. D., Bombaci, S., Caravaggi, A., Chalmers, A., Clulow, S., Griffin, A., Hayward, M., Heurich, M., Linnell, J., Marnewick, K., Moehrenschlager, A., Montgomery, A., Shuttleworth, C., Somers, M. & Weise, F., Nov 2019, In: Science. 364, p. 6438Research output: Contribution to journal › Letter
- Published
Pine marten scat holds few clues for squirrel disease
Shuttleworth, C., Everest, D. J., Dastjerdi, A., Birks, J. & Hayward, M., 7 Dec 2019, In: Veterinary Record. 185, p. 698Research output: Contribution to journal › Article
- 2020
- Published
Invasive alien species colonisation prevention: Your guide to early detection and rapid response
Robinson, N. & Shuttleworth, C., 2020, Newark: Royal Society of Wildlife Trusts. 204 p.Research output: Book/Report › Book › peer-review
- Published
Envisioning the future with 'compassionate conservation': An ominous projection for native wildlife and biodiversity
Callen, A., Hayward, M. W., Klop-Toker, K., Allen, B. L., Ballard, G., Beranek, C. T., Broekhuis, F., Bugir, C. K., Clarke, R. H., Clulow, J., Clulow, S., Daltry, J. C., Davies-Mostert, H. T., Di Blanco, Y. E., Dixon, V., Fleming, P. J. S., Howell, L. G., Kerley, G. I. H., Legge, S. M., Lenga, D. J., Major, T., Montgomery, R. A., Moseby, K., Meyer, N., Parker, D. M., Periquet, S., Read, J., Scanlon, R. J., Shuttleworth, C., Tamessar, C. T., Taylor, W. A., Tuft, K., Upton, R. M. O., Valenzuela, M., Witt, R. R. & Wuster, W., Jan 2020, In: Biological Conservation. 241, 108365.Research output: Contribution to journal › Article › peer-review
- Published
Response to comments on "Compassionate Conservation deserves a morally serious rather than dismissive response - reply to Callen et al., 2020"
Callen, A., Hayward, M. W., Klop-Toker, K., Allen, B. L., Ballard, G., Beranek, C. T., Broekhuis, F., Bugir, C. K., Clarke, R. H., Clulow, J., Clulow, S., Daltry, J. C., Davies-Mostert, H. T., Di Blanco, Y. E., Dixon, V., Fleming, P. J. S., Howell, L. G., Kerley, G. I. H., Legge, S. M., Lenga, D. J., Major, T., Montgomery, R. A., Moseby, K., Meyer, N., Parker, D. M., Periquet, S., Read, J., Scanlon, R. J., Shuttleworth, C., Tamessar, C. T., Taylor, W. A., Tuft, K., Upton, R. M. O., Valenzuela, M., Witt, R. R. & Wuster, W., 30 Apr 2020, In: Biological Conservation. 244Research output: Contribution to journal › Letter › peer-review
- Published
Introduced Canadian Eastern grey squirrels: squirrelpox virus surveillance and why nothing matters.
McInnes, C., Shuttleworth, C., Larsen, K. W., Everest, D. J., Bruemmer, C., Carroll, B., Romeo, C., Sainsbury, A. W., Crawshaw, G., Dubois, S., Gillis, L., Gilray, J. & Percival, A., 3 Aug 2020, In: Hystrix-Italian Journal of Mammalogy. 31, p. 95-98Research output: Contribution to journal › Article › peer-review
- Published
Challenges and solutions in early detection, rapid response and communication about potential invasive alien species in forests
de Groot, M., Bullas-Appleton, E., Csóka, G., Csiszár, A., Faccoli, A., Gervasini, A., Kirichenko, N., Shuttleworth, C., Kus Veenvliet, J., O'Hanlon, R., Korda, M., Marinsek, A., Robinson, N., Sweeney, J., Tricarico, E., Verbrugge, L., Williams, D. & Zidar, S., 14 Aug 2020, In: Management of Biological Invasions. 11, p. 637-660Research output: Contribution to journal › Article › peer-review
- Published
Do differing levels of boldness influence the success of translocation? A pilot study on red squirrels (Sciurus vulgaris)
Bamber, J. A., Shuttleworth, C. & Hayward, M., 26 Sept 2020, In: Animals. 10, 1748.Research output: Contribution to journal › Article › peer-review
- Published
Evolving grey squirrel management techniques in Europe
Shuttleworth, C., Robinson, N. J., Halliwell, E. C., Clews-Roberts, R., Peek, H., Podgornik, G., Stinson, M., Rice, S., Finlay, C., McKinney, C., Everest, D. J. & Larsen, K. W., 21 Oct 2020, In: Management of Biological Invasions. 11, 4, p. 747-761Research output: Contribution to journal › Article › peer-review
- Published
Are novel ecosystems the only novelty of rewilding?
Klop-Toker, K., Clulow, S., Shuttleworth, C. & Hayward, M., 1 Nov 2020, In: Restoration Ecology. 28, p. 1318-1320Research output: Contribution to journal › Article › peer-review
- Published
Ethical dilemmas when using citizen science for early detection of invasive tree pests and diseases
Pocock, M. J. O., Marzano, M., Bullas-Appleton, E., Dyke, A., de Groot, M., Shuttleworth, C. & White, R., 1 Nov 2020, In: Management of Biological Invasions. 11, p. 720-732Research output: Contribution to journal › Article › peer-review
- Published
Reinstating trophic cascades as an applied conservation tool to protect forest ecosystems from invasive grey squirrels (Sciurus carolinensis)
Bamber, J. A., Shuttleworth, C. M., Hayward, M. W. & Everest, D. J., 31 Dec 2020, In: Food Webs. 25, e00164.Research output: Contribution to journal › Article › peer-review
- 2021
- Published
A practical guide to early surveillance and monitoring
Robinson, N. J., Everest, D. J., Harry, P. & Shuttleworth, C., 2021, Invasive Alien Species Colonisation Prevention: Your guide to early detection and rapid response . Robinson, N. J. & Shuttleworth, C. M. (eds.). Royal Society of Wildlife Trusts, p. 163-202Research output: Chapter in Book/Report/Conference proceeding › Chapter
- Published
Magical Mammals: Red squirrels in North West Wales
Shuttleworth, C., 2021, Saving the red squirrel: landscape recovery. Shuttleworth, C. M., Lurz, P. W. W. & Robinson, N. J. (eds.). Red Squirrel Survival Trust, p. 269-278Research output: Chapter in Book/Report/Conference proceeding › Chapter
- Published
Monitoring escaped captive bred red squirrels in the UK
Shuttleworth, C., 2021, Saving the red squirrel: landscape recovery. Shuttleworth, C. M., Lurz, P. W. W. & Robinson, N. J. (eds.). Red Squirrel Survival Trust, p. 71-78Research output: Chapter in Book/Report/Conference proceeding › Chapter
- Published
Preventing grey squirrel establishment on Anglesey
Shuttleworth, C., 2021, Invasive Alien Species Colonisation Prevention: Your guide to early detection and rapid response . Robinson, N. J. & Shuttleworth, C. M. (eds.). Royal Society of Wildlife Trusts, p. 55-69Research output: Chapter in Book/Report/Conference proceeding › Chapter
- Published
Red Squirrel Conservation in Clocaenog Forest: Review and Recommendations
Shuttleworth, C., 2021, 56 p.Research output: Book/Report › Commissioned report