Dr Craig Shuttleworth
Honorary Visiting Research Fellow

Affiliations
Links
- https://twitter.com/Welshsquirrels
Personal account - https://twitter.com/RedSquirrelsinW
Welsh red squirrel conservation - https://www.facebook.com/groups/143610382316300
Welsh red squirrel conservation - https://pinemarten.net/
The Gwynedd pine marten project - https://www.linkedin.com/in/craig-shuttleworth-6062a1253/
Personal page
Contact info
Mobile 07966150847
Email c.shuttleworth@bangor.ac.uk
- Published
A practical guide to early surveillance and monitoring
Robinson, N. J., Everest, D. J., Harry, P. & Shuttleworth, C., 2021, Invasive Alien Species Colonisation Prevention: Your guide to early detection and rapid response . Robinson, N. J. & Shuttleworth, C. M. (eds.). Royal Society of Wildlife Trusts, p. 163-202Research output: Chapter in Book/Report/Conference proceeding › Chapter
- Published
A preliminary comparison of magnum 116 body-grip traps with a live capture trap design during the control of grey squirrels (Sciurus carolinensis)
Shuttleworth, C., Mill, A. & Van der Waal, Z., Dec 2016, In: International Journal of Pest Management. 63, 4, p. 316-321Research output: Contribution to journal › Article › peer-review
- Published
A study of nest box use by a low density red squirrel (Sciurus vulgaris) population within a commercial conifer plantation
Shuttleworth, C., 2004, In: Quarterly Journal of Forestry. 98, p. 113-119Research output: Contribution to journal › Article › peer-review
- Published
A systematic assessment of the impact of adenovirus infection on a captive re-introduction project for red squirrels (Sciurus vulgaris)
Everest, D. J., Shuttleworth, C., Grierson, S. S., Duff, J. P., Jackson, N., Litherland, P. & Kenward, R. E., 31 Aug 2012, In: Veterinary Record. 171, p. 176-184Research output: Contribution to journal › Article › peer-review
- Published
A temporal refuge from predation can change the outcome of prey species competition
Slade, A., White, A., Lurz, P. W. W., Shuttleworth, C. & Lambin, X., 1 Sept 2022, In: Oikos. Oikos e08565Research output: Contribution to journal › Article › peer-review
- Published
A tough nut to crack: red squirrel conservation in Wales
Shuttleworth, C., 1 Oct 2003, In: Biologist. 50, p. 231-235Research output: Contribution to journal › Article › peer-review
- Published
Adenovirus detections in juvenile squirrels
Everest, D. J., Dastjerdi, A., Cowan, D., Gomm, M., Shuttleworth, C., McInnes, C. J., Deane, D., Coulter, L., Mill, A., Rushton, S. P., Jackson, N. L. & Litherland, P., 1 Oct 2015, In: Veterinary Record. 177, p. 373-374Research output: Contribution to journal › Article
- Published
Adenovirus infection in red squirrels
Shuttleworth, C., Everest, D. J., Stidworthy, M., Halliwell, E. C., Clews-Roberts, R., Paine, V., Bamber, C. & Mason, J., 31 Dec 2021, In: The Veterinary record. 189, p. 444-445Research output: Contribution to journal › Article
- Published
Adenovirus infection in red squirrels in areas free from grey squirrels
Everest, D. J., Butler, H., Simpson, V. R., Blackett, T. & Shuttleworth, C., 31 Aug 2013, In: The Veterinary record. 173, 8, p. 199-200Research output: Contribution to journal › Article
- Published
Adenovirus infection: An emerging factor in red squirrel conservation programmes
Everest, D. J., Shuttleworth, C., Stidworthy, M. F., Grierson, S. S., Duff, P. J. & Kenward, R. E., 1 Oct 2014, In: Mammal Review. 44, 3-4, p. 225-233Research output: Contribution to journal › Article › peer-review
- Published
Adenovirus remains widespread in squirrels on Anglesey
Shuttleworth, C., Everest, D. & McQueen, A., 16 Dec 2023, In: Veterinary Record. 193, 12, p. 490-491 2 p.Research output: Contribution to journal › Letter
- Published
Adenovirus-associated deaths in red squirrels on Anglesey
Everest, D. J., Shuttleworth, C. & Stidworthy, M. F., 31 Oct 2008, In: The Veterinary record. 163, 14, p. 430Research output: Contribution to journal › Article › peer-review
- Published
Against a grey tide: What future for the red squirrel?
Shuttleworth, C., 2005, Journal of Practical Ecology & Conservation (Special Series No 4): Living with the Aliens? - Ecology, History, Culture and Management if exotic plants and animals: issues for nature conservation . Rotherham, L. D. (ed.). Vol. 4. p. 87-88Research output: Chapter in Book/Report/Conference proceeding › Chapter
- Published
Against a grey tide: red squirrel conservation on Anglesey
Shuttleworth, C., 2005, In: Quarterly Journal of Forestry. 99, p. 213-214Research output: Contribution to journal › Article › peer-review
- Published
An Opportunistic Assessment of the Impact of Squirrelpox Disease Outbreaks upon a Red Squirrel Population Sympatric with Grey Squirrels in Wales
Shuttleworth, C., Everest, D., Holmes, P., Bell, S. & Cripps, R., 1 Jan 2022, In: Animals. 12, 1Research output: Contribution to journal › Article › peer-review
- Published
Are nest boxes a useful tool in regional red squirrel conservation programs?
Shuttleworth, C. & Schuchert , P., 15 Dec 2014, In: Hystrix-Italian Journal of Mammalogy. 25, p. 91-94Research output: Contribution to journal › Article › peer-review
- Published
Are novel ecosystems the only novelty of rewilding?
Klop-Toker, K., Clulow, S., Shuttleworth, C. & Hayward, M., 1 Nov 2020, In: Restoration Ecology. 28, p. 1318-1320Research output: Contribution to journal › Article › peer-review
- Published
Assessing a potential non-invasive method for viral diagnostic purposes in European squirrels
Everest, D. J., Tolhurst-Cherriman, D. A. R., Davies, H., Dastjerdi, A., Ashton, A., Blackett, T., Meredith, A. L., Milne, E., Mill, A. & Shuttleworth, C., 4 Jun 2019, In: Hystrix-Italian Journal of Mammalogy. 30, p. 44-50Research output: Contribution to journal › Article › peer-review
- Published
Assessing causes and significance of red squirrel (Sciurus vulgaris) mortality during regional population restoration: An applied conservation perspective
Shuttleworth, C., Signorile, , A. L., Everest, D. J., Lurz, P. W. W. & Duff, J. P., 16 Nov 2015, In: Hystrix-Italian Journal of Mammalogy. 26, 2, p. 69-75Research output: Contribution to journal › Article › peer-review
- Published
British red squirrel strongholds: challenges for conservation
Gurnell, J., Blackett, T., Butler, H., Lurz, P. W. W., Magris, L. & Shuttleworth, C., 2015, Red squirrels: Ecology, Conservation & Management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Hayward, M. W. (eds.). European Squirrel Initiative, p. 211-232Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review
- Published
British red squirrels remain the only known wild rodent host for leprosy bacilli
Schilling, A., Avanzi, C., Ulrich, R. G., Busso, P., Pisanu, B., Ferrari, N., Romeo, C., Mazzamuto, M. V., McLuckie, J., Shuttleworth, C., Del-Pozo, J., Lurz, P. W. W., Escalante-Fuentes, W. G., Ocampo-Candiani, J., Vera-Cabrera, L., Stevenson, K., Chapuis, J.-L., Meredith, A. L. & Cole, S. T., 1 Feb 2019, In: Frontiers in Veterinary Science. 6, p. 8Research output: Contribution to journal › Article › peer-review
- Published
By trying to be everything, rewilding risks meaning nothing
Jachowski, J. D., Bombaci, S., Caravaggi, A., Chalmers, A., Clulow, S., Griffin, A., Hayward, M., Heurich, M., Linnell, J., Marnewick, K., Moehrenschlager, A., Montgomery, A., Shuttleworth, C., Somers, M. & Weise, F., Nov 2019, In: Science. 364, p. 6438Research output: Contribution to journal › Letter
- Published
Capturing red squirrels (Sciurus vulgaris) on camera: A cost-effective approach for monitoring relative abundance and habitat preference
Shannon, G., Valle, S. & Shuttleworth, C., 3 Oct 2023, In: Ecology and Evolution. 13, 10, e10536.Research output: Contribution to journal › Article › peer-review
- Published
Challenges and solutions in early detection, rapid response and communication about potential invasive alien species in forests
de Groot, M., Bullas-Appleton, E., Csóka, G., Csiszár, A., Faccoli, A., Gervasini, A., Kirichenko, N., Shuttleworth, C., Kus Veenvliet, J., O'Hanlon, R., Korda, M., Marinsek, A., Robinson, N., Sweeney, J., Tricarico, E., Verbrugge, L., Williams, D. & Zidar, S., 14 Aug 2020, In: Management of Biological Invasions. 11, p. 637-660Research output: Contribution to journal › Article › peer-review
- Published
Confirmation of squirrelpox in Welsh red squirrel
Everest, D. J., Floyd, T., Donnnacie, B., Irvine, R. M., Holmes, J. P. & Shuttleworth, C., 30 Nov 2017, In: Veterinary Record. 181, p. 514-515Research output: Contribution to journal › Article
- Published
Controlling grey squirrels in forests and woodlands in the UK
Gill, R., Ferryman, M., Shuttleworth, C., Lurz, P. W. W., Mill, A., Robertson, P. A. & Dutton, C., 2019, Forest Research. 16 p.Research output: Book/Report › Book › peer-review
- Published
Deconstructing compassionate conservation
Hayward, M. W., Callen, A., Allen, B. L., Ballard, G., Broekhuis, F., Bugir, C., Clarke, R. H., Clulow, J., Clulow, S., Daltry, J. C., Davies-Mostert, H. T., Fleming, P. J. S., Griffin, A. S., Howell, L. G., Kerley, G. I. H., Klop-Toker, K., Legge, S., Major, T., Meyer, N., Montgomery, R. A., Moseby, K., Parker, D. M., Périquet, S., Read, J., Scanlon, R., Seeto, R., Shuttleworth, C., Somers, M. J., Tamessar, C. T., Tuft, K., Upton, R., Valenzuela-Molina, M., Wayne, A., Witt, R. R. & Wuster, W., Aug 2019, In: Conservation Biology. 33, 4, p. 760-768Research output: Contribution to journal › Article › peer-review
- Published
Detecting viral infection in red squirrels
Shuttleworth, C., Everest, D. J., Halliwell, E. C., Hulme, B., Wilberforce, L. & Clews-Roberts, R., 30 Apr 2019, In: Veterinary Record. 184, p. 507Research output: Contribution to journal › Article
- Published
Developing a method for assessing the relative abundance of red squirrels in low-density populations
Shannon, G., Barton, O., Valle, S. & Shuttleworth, C., 1 Nov 2024, 34 p.Research output: Book/Report › Commissioned report
- Published
Developing integrated and applied red squirrel conservation programmes: What lessons can Europe learn from a regional grey squirrel eradication programme in North Wales?
Shuttleworth, C., Schuchert, P., Everest, D. J., McInnes, C. J., Rushton, S. P., Jackson, N. & Kenward, R. E., 2015, Red squirrels: Ecology, Conservation & Management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Hayward, M. W. (eds.). European Squirrel Initiative, p. 233-250Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review
- Published
Developing red squirrel re-introduction techniques for use during regional grey squirrel eradication programmes in Europe
Shuttleworth, C., Jackson, N. & Kenward, R. E., 2015, 2015 Global Reintroduction Perspectives. Soorae, P. (ed.). IUCN/SSC Re-introduction Specialist Group, p. 67-74Research output: Chapter in Book/Report/Conference proceeding › Chapter
- Published
Disease monitoring and surveillance: case studies in the applied conservation of fragmented red squirrel (Sciurus vulgaris) populations in England and Wales
Everest, D. J., Floyd, T., Holmes, P., Man, C., Dunnett, C., Locke, R., Savage, L., Sutcliffe, S., Sapsford, B., Shuttleworth, C. & Duff, P., 1 Dec 2021, In: Mammalian Biology. 101, p. 1079-1088Research output: Contribution to journal › Article › peer-review
- Published
Disease, invasions and conservation: no evidence of squirrelpox virus in grey squirrels introduced to Italy
Romeo, C., McInnes, C., Dale, T., Shuttleworth, C., Bertolino, S., Wauters, L. A. & Ferrari, N., 1 Feb 2019, In: Animal Conservation. 22, 1, p. 14-23Research output: Contribution to journal › Article › peer-review
- Published
Do differing levels of boldness influence the success of translocation? A pilot study on red squirrels (Sciurus vulgaris)
Bamber, J. A., Shuttleworth, C. & Hayward, M., 26 Sept 2020, In: Animals. 10, 1748.Research output: Contribution to journal › Article › peer-review
- Published
Ecosystem impacts of an alien invader in Europe, the grey squirrel Sciurus carolinensis
Gurnell, J., Lurz, P. W. W. & Shuttleworth, C., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (eds.). European Squirrel Initiative, p. 307-329Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review
- Published
Envisioning the future with 'compassionate conservation': An ominous projection for native wildlife and biodiversity
Callen, A., Hayward, M. W., Klop-Toker, K., Allen, B. L., Ballard, G., Beranek, C. T., Broekhuis, F., Bugir, C. K., Clarke, R. H., Clulow, J., Clulow, S., Daltry, J. C., Davies-Mostert, H. T., Di Blanco, Y. E., Dixon, V., Fleming, P. J. S., Howell, L. G., Kerley, G. I. H., Legge, S. M., Lenga, D. J., Major, T., Montgomery, R. A., Moseby, K., Meyer, N., Parker, D. M., Periquet, S., Read, J., Scanlon, R. J., Shuttleworth, C., Tamessar, C. T., Taylor, W. A., Tuft, K., Upton, R. M. O., Valenzuela, M., Witt, R. R. & Wuster, W., Jan 2020, In: Biological Conservation. 241, 108365.Research output: Contribution to journal › Article › peer-review
- Published
Ethical dilemmas when using citizen science for early detection of invasive tree pests and diseases
Pocock, M. J. O., Marzano, M., Bullas-Appleton, E., Dyke, A., de Groot, M., Shuttleworth, C. & White, R., 1 Nov 2020, In: Management of Biological Invasions. 11, p. 720-732Research output: Contribution to journal › Article › peer-review
- Published
Evolving grey squirrel management techniques in Europe
Shuttleworth, C., Robinson, N. J., Halliwell, E. C., Clews-Roberts, R., Peek, H., Podgornik, G., Stinson, M., Rice, S., Finlay, C., McKinney, C., Everest, D. J. & Larsen, K. W., 21 Oct 2020, In: Management of Biological Invasions. 11, 4, p. 747-761Research output: Contribution to journal › Article › peer-review
- Published
Experiences of grey squirrel management in an upland conifer forest
Halliwell, E. C., Shuttleworth, C., Cartmel, S., Lloyd, I. & Jenkins, R., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (eds.). European Squirrel Initiative, p. 453-474Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review
- Published
First report of dermatophilosis in wild European red squirrels (Sciurus vulgaris)
Holmes, P., Everest, D. J., Spiro, S., Wessels, M. & Shuttleworth, C., 30 Sept 2019, In: Veterinary Record Case Reports. 7, 3, p. 3Research output: Contribution to journal › Article › peer-review
- Published
Future red squirrel conservation in Clocaenog forest
Shuttleworth, C. & Hayward, M., 2015, P21018-0026 ed.Research output: Book/Report › Commissioned report
- Published
Genetic evidence of the effectiveness of grey squirrel control operations: lessons from the Isle of Anglesey
Signorile, L. & Shuttleworth, C., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (eds.). European Squirrel Initiative, p. 439-452Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review
- Published
Genetic management of the red squirrel, Sciurus vulgaris: a practical approach to regional conservation
Ogden, R., Shuttleworth, C., McEwing, R. & Cesarini, S., 26 Oct 2005, In: Conservation Genetics. 6, p. 511-525Research output: Contribution to journal › Article › peer-review
- Published
Grey squirrel control as a tactic in the conservation of the red squirrel: a positive outcome
Shuttleworth, C., 2001, Advances in Vertebrate Pest Management . Feare, C. J. & Cowan, D. P. (eds.). Filander Verlag, Vol. 3. p. 151-168Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review
- Published
Grey squirrel management in Europe - the future
Shuttleworth, C., Lurz, P. W. W. & Gurnell, J., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (eds.). European Squirrel InitiativeResearch output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review
- Published
Grey squirrel nesting ecology and the use of nest sites in European population management
Shuttleworth, C., Selonen, V. & Koprowski, J. L., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (eds.). European Squirrel Initiative, p. 349-369Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review
- Published
Grey squirrels are bad for the British countryside – full stop
Shuttleworth, C., 2017, The Conversation.Research output: Other contribution
- Published
Gut bacteria may explain why grey squirrels outcompete reds – new research
Shuttleworth, C., 20 Feb 2024, The Conversation.Research output: Contribution to specialist publication › Article
- Published
Identifying incursion pathways, early detection responses and management actions to prevent grey squirrel range expansion: an island case study in Wales
Shuttleworth, C., Halliwell, E. C. & Robertson, P. A., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (eds.). European Squirrel Initiative, p. 475-494Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review
- Published
Indirect effects of pine marten recovery result in benefits to native prey through suppression of an invasive species and a shared pathogen
Slade, A., White, A., Lurz, P. W. W., Shuttleworth, C., Tosh, D. & Twining, J., 1 Feb 2023, In: Ecological Modelling. 476, 110216.Research output: Contribution to journal › Article › peer-review