Dr Craig Shuttleworth
Honorary Visiting Research Fellow

Affiliations
Links
- https://twitter.com/Welshsquirrels
Personal account - https://twitter.com/RedSquirrelsinW
Welsh red squirrel conservation - https://www.facebook.com/groups/143610382316300
Welsh red squirrel conservation - https://pinemarten.net/
The Gwynedd pine marten project - https://www.linkedin.com/in/craig-shuttleworth-6062a1253/
Personal page
Contact info
Mobile 07966150847
Email c.shuttleworth@bangor.ac.uk
- 2019
- Published
Why the pine marten is not every red squirrel’s best friend
Shuttleworth, C. & Hayward, M., 2019, The Conversation.Research output: Other contribution
- 2018
- Published
Reducing negative impacts on non-target species during grey squirrel (Sciurus carolinensis) control activities
Shuttleworth, C., Mill, A. C., Van der Wall, Z., Bertolino, S. & Everest, D. J., 1 Jan 2018, In: Quarterly Journal of Forestry. 112, 1, p. 21-29Research output: Contribution to journal › Article › peer-review
- Published
The implications of significant adenovirus infection in UK captive red squirrel (Sciurus vulgaris) collections: How histological screening can aid applied conservation management
Everest, D. J., Shuttleworth, C., Grierson, S. S., Dastjerdi, A., M.F., S., Duff, J. P., Higgins, R. J., Mill, A. C. & Chantrey, J., 1 Jan 2018, In: Mammalian Biology. 88, p. 123-129Research output: Contribution to journal › Article › peer-review
- Published
Rare woodland wildlife at risk because of 50-year-old tree felling rules
Shuttleworth, C., 2018, The Conversation.Research output: Other contribution
- Published
Red squirrel (Sciurus vulgaris) conservation plan for Anglesey & Gwynedd
Shuttleworth, C. & Halliwell, E. C., 2018, 50 p.Research output: Book/Report › Commissioned report
- Published
Red squirrels in my garden: guidance and tips to help conserve local populations
Shuttleworth, C. & Halliwell, E. C., 2018, European Squirrel Initiative. 185 p.Research output: Book/Report › Book › peer-review
- Published
Wales should follow Scotland’s lead in protecting the red squirrel’s habitat
Shuttleworth, C., 2018, Nation Cymru .Research output: Other contribution
- 2017
- Published
Confirmation of squirrelpox in Welsh red squirrel
Everest, D. J., Floyd, T., Donnnacie, B., Irvine, R. M., Holmes, J. P. & Shuttleworth, C., 30 Nov 2017, In: Veterinary Record. 181, p. 514-515Research output: Contribution to journal › Article
- Published
Mathematical models for invasive species management: grey squirrel control on Anglesey
Jones, H., White, A., Lurz, P. W. W. & Shuttleworth, C., 10 Sept 2017, In: Ecological Modelling. 180, p. 274-284Research output: Contribution to journal › Article › peer-review
- Published
Virus presence within Welsh red squirrels
Everest, D. J., Holmes, P. & Shuttleworth, C., 30 May 2017, In: The Veterinary record. 180, p. 520-521Research output: Contribution to journal › Article
- Published
The large-scale removal of mammalian invasive alien species in Northern Europe
Robertson, P. A., Adriaens, T., Lambin, X., Mill, A., Roy, S., Shuttleworth, C. & Sutton-Croft, M., 1 Feb 2017, In: Pest Management Science. 73, p. 273-279Research output: Contribution to journal › Article › peer-review
- Published
Grey squirrels are bad for the British countryside – full stop
Shuttleworth, C., 2017, The Conversation.Research output: Other contribution
- 2016
- Published
A preliminary comparison of magnum 116 body-grip traps with a live capture trap design during the control of grey squirrels (Sciurus carolinensis)
Shuttleworth, C., Mill, A. & Van der Waal, Z., Dec 2016, In: International Journal of Pest Management. 63, 4, p. 316-321Research output: Contribution to journal › Article › peer-review
- Published
Ecosystem impacts of an alien invader in Europe, the grey squirrel Sciurus carolinensis
Gurnell, J., Lurz, P. W. W. & Shuttleworth, C., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (eds.). European Squirrel Initiative, p. 307-329Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review
- Published
Experiences of grey squirrel management in an upland conifer forest
Halliwell, E. C., Shuttleworth, C., Cartmel, S., Lloyd, I. & Jenkins, R., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (eds.). European Squirrel Initiative, p. 453-474Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review
- Published
Genetic evidence of the effectiveness of grey squirrel control operations: lessons from the Isle of Anglesey
Signorile, L. & Shuttleworth, C., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (eds.). European Squirrel Initiative, p. 439-452Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review
- Published
Grey squirrel management in Europe - the future
Shuttleworth, C., Lurz, P. W. W. & Gurnell, J., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (eds.). European Squirrel InitiativeResearch output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review
- Published
Grey squirrel nesting ecology and the use of nest sites in European population management
Shuttleworth, C., Selonen, V. & Koprowski, J. L., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (eds.). European Squirrel Initiative, p. 349-369Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review
- Published
Identifying incursion pathways, early detection responses and management actions to prevent grey squirrel range expansion: an island case study in Wales
Shuttleworth, C., Halliwell, E. C. & Robertson, P. A., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (eds.). European Squirrel Initiative, p. 475-494Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review
- Published
Mathematical models of grey squirrel invasion: a case study on Anglesey
Jones, H. E., White, A., Lurz, P. W. W., Boots, M. & Shuttleworth, C., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (eds.). European Squirrel Initiative, p. 235-255Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review
- Published
Ranging behaviour, density and social structure in grey squirrels
Lawton, C., Shuttleworth, C. & Kenward, R. E., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (eds.). European Squirrel Initiative, p. 133-152Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review
- Published
The grey squirrel: ecology & management of an Invasive Species in Europe
Shuttleworth, C., Lurz, P. W. W. & Gurnell, J., 2016, European Squirrel Initiative. 505 p.Research output: Book/Report › Book › peer-review
- Published
The management of grey squirrel populations in Europe: evolving best practice
Bertolino, S., Lurz, P. W. W., Shuttleworth, C., Martinoli, A. & Wauters, L. A., 2016, The grey squirrel: ecology and management in Europe . Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (eds.). European Squirrel Initiative, p. 495-516Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review
- Published
The potential role of ectoparasites in the epidemiology of squirrelpox virus: a possible novel means of intervention to reduce the impact of the disease?
Cowan, D. P., Mill, A. C., Everest, D. J., McInnes, C. J., Rushton, S. P., Shirley, M. D. F., Start, C. & Shuttleworth, C., 2016, The grey squirrel: ecology and management in Europe . Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (eds.). European Squirrel Initiative, p. 255-289Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review
- 2015
- Published
Assessing causes and significance of red squirrel (Sciurus vulgaris) mortality during regional population restoration: An applied conservation perspective
Shuttleworth, C., Signorile, , A. L., Everest, D. J., Lurz, P. W. W. & Duff, J. P., 16 Nov 2015, In: Hystrix-Italian Journal of Mammalogy. 26, 2, p. 69-75Research output: Contribution to journal › Article › peer-review
- Published
Adenovirus detections in juvenile squirrels
Everest, D. J., Dastjerdi, A., Cowan, D., Gomm, M., Shuttleworth, C., McInnes, C. J., Deane, D., Coulter, L., Mill, A., Rushton, S. P., Jackson, N. L. & Litherland, P., 1 Oct 2015, In: Veterinary Record. 177, p. 373-374Research output: Contribution to journal › Article
- Published
British red squirrel strongholds: challenges for conservation
Gurnell, J., Blackett, T., Butler, H., Lurz, P. W. W., Magris, L. & Shuttleworth, C., 2015, Red squirrels: Ecology, Conservation & Management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Hayward, M. W. (eds.). European Squirrel Initiative, p. 211-232Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review
- Published
Developing integrated and applied red squirrel conservation programmes: What lessons can Europe learn from a regional grey squirrel eradication programme in North Wales?
Shuttleworth, C., Schuchert, P., Everest, D. J., McInnes, C. J., Rushton, S. P., Jackson, N. & Kenward, R. E., 2015, Red squirrels: Ecology, Conservation & Management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Hayward, M. W. (eds.). European Squirrel Initiative, p. 233-250Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review
- Published
Developing red squirrel re-introduction techniques for use during regional grey squirrel eradication programmes in Europe
Shuttleworth, C., Jackson, N. & Kenward, R. E., 2015, 2015 Global Reintroduction Perspectives. Soorae, P. (ed.). IUCN/SSC Re-introduction Specialist Group, p. 67-74Research output: Chapter in Book/Report/Conference proceeding › Chapter
- Published
Future red squirrel conservation in Clocaenog forest
Shuttleworth, C. & Hayward, M., 2015, P21018-0026 ed.Research output: Book/Report › Commissioned report
- Published
Island Haven: Anglesey red squirrel conservation
Shuttleworth, C., 2015, Shared Experience of Red Squirrel Conservation Practice . Shuttleworth, C. M., Lurz, P. W. W. & Halliwell, E. C. (eds.). European Squirrel Initiative, p. 151-158Research output: Chapter in Book/Report/Conference proceeding › Chapter
- Published
Painting the Town Red: Gwynedd red squirrel conservation
Shuttleworth, C., 2015, Shared Experience of Red Squirrel Conservation Practice . Shuttleworth, C. M., Lurz, P. W. W. & Halliwell, E. C. (eds.). European Squirrel Initiative, p. 159-166Research output: Chapter in Book/Report/Conference proceeding › Chapter
- Published
Red squirrels: ecology, conservation & management in Europe
Shuttleworth, C., Hayward, M. & Lurz, P. W. W., 2015, European Squirrel Initiative. 330 p.Research output: Book/Report › Book › peer-review
- Published
Reintroductions and translocations of red squirrels within Europe
Lawton, C., Waters, C. & Shuttleworth, C., 2015, Red squirrels: Ecology, Conservation & Management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Hayward, M. W. (eds.). European Squirrel Initiative, p. 193-210Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review
- Published
Shared experience of red squirrel conservation practice
Shuttleworth, C., Lurz, P. W. W. & Halliwell, E. C., 2015, European Squirrel Initiative. 220 p.Research output: Book/Report › Book
- Published
Striving for success: an evaluation of local action to conserve red squirrels (Sciurus vulgaris) in Wales
Halliwell, E. C., Shuttleworth, C., Wilberforce, L., Denman, H., Lloyd, I. & Cartmel, S., 2015, Red squirrels: Ecology, Conservation & Management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Hayward, M. W. (eds.). European Squirrel Initiative, p. 175-192Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review
- Published
The impact of adenoviruses on red squirrel (Sciurus vulgaris) conservation – with notes on other enteric viruses of potential significance
Everest, D. J., Shuttleworth, C., Grierson, S. S., Dastjerdi, A., Stidworthy, M. F., Duff, J. P. & Sainsbury, A. W., 2015, Red squirrels: Ecology, Conservation & Management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Hayward, M. W. (eds.). European Squirrel Initiative, p. 129-146Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review
- 2014
- Published
Are nest boxes a useful tool in regional red squirrel conservation programs?
Shuttleworth, C. & Schuchert , P., 15 Dec 2014, In: Hystrix-Italian Journal of Mammalogy. 25, p. 91-94Research output: Contribution to journal › Article › peer-review
- Published
Landscape scale impacts of culling upon a European grey squirrel population: can trapping reduce population size and decrease the threat of squirrel pox virus infection for the native red squirrel?
Schuchert, P., Shuttleworth, C., McInnes, C. J., Everest, D. J. & Rushton, S. P., 1 Nov 2014, In: Biological Invasions. 16, p. 2381-2391Research output: Contribution to journal › Article › peer-review
- Published
Adenovirus infection: An emerging factor in red squirrel conservation programmes
Everest, D. J., Shuttleworth, C., Stidworthy, M. F., Grierson, S. S., Duff, P. J. & Kenward, R. E., 1 Oct 2014, In: Mammal Review. 44, 3-4, p. 225-233Research output: Contribution to journal › Article › peer-review
- Published
Inter-specific viral infections: Can the management of captive red squirrel collections help inform scientific research?
Shuttleworth, C., Everest, D. J., McInnes, C. J., Greenwood, A., Jackson, N., Rushton, S. P. & Kenward, R. E., 16 Jun 2014, In: Hystrix-Italian Journal of Mammalogy. 25, 1, p. 18-24Research output: Contribution to journal › Article › peer-review
- Published
Red squirrel conservation and disease - Adenovirus in Wales
Shuttleworth, C., 2014, In: Natur Cymru. 52, p. 41-44Research output: Contribution to journal › Article
- Published
The role of spruce in red squirrel (Sciurus vulgaris) conservation in the UK
Shuttleworth, C. & Lurz, P. W. W., 2014, Forest Ecosystems: Biodiversity, Management and Conservation. Roberts, N. C. (ed.). Nova Science Publishers, p. 63-78Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review
- 2013
- Published
Adenovirus infection in red squirrels in areas free from grey squirrels
Everest, D. J., Butler, H., Simpson, V. R., Blackett, T. & Shuttleworth, C., 31 Aug 2013, In: The Veterinary record. 173, 8, p. 199-200Research output: Contribution to journal › Article
- 2012
- Published
Integrating red squirrel (Sciurus vulgaris) habitat requirements with the management of pathogenic tree disease in commercial forests in the UK
Shuttleworth, C., Lurz, P. W. W., Geddes, N. & Brown, J., 1 Sept 2012, In: Forest Ecology and Management. 279, p. 167-175Research output: Contribution to journal › Article › peer-review
- Published
A systematic assessment of the impact of adenovirus infection on a captive re-introduction project for red squirrels (Sciurus vulgaris)
Everest, D. J., Shuttleworth, C., Grierson, S. S., Duff, J. P., Jackson, N., Litherland, P. & Kenward, R. E., 31 Aug 2012, In: Veterinary Record. 171, p. 176-184Research output: Contribution to journal › Article › peer-review
- Published
Red squirrel population monitoring: Clocaenog forest
Shuttleworth, C. & Donoyou, T., 2012, IFCW108/001/002 ed.Research output: Book/Report › Commissioned report
- 2010
-
Retrospective detection by negative contrast electron microscopy of faecal viral particles in wild red squirrels (Sciurus vulgaris) with suspected enteropathy in Great Britain
Everest, D. J., Stidworthy, M. F., Milne, E. M., Meredith, A. L., Chantery, J., Shuttleworth, C., Blackett, T., Butler, H., Wilkinson, M. & Sainsbury, A. W., 31 Dec 2010, In: Veterinary Record. 167, p. 1007-1010Research output: Contribution to journal › Article › peer-review
- Published
Turning the grey tide: progress in red squirrel recovery
Shuttleworth, C., 1 Jun 2010, In: Ecos. 31, p. 27-35Research output: Contribution to journal › Article
- 2009
- Published
PCR detection of adenovirus in grey squirrels on Anglesey
Everest, D. J., Grierson, S. S., Stidworthy, M. & Shuttleworth, C., 31 Oct 2009, In: Veterinary Record. 165, p. 482Research output: Contribution to journal › Article › peer-review