Dr Craig Shuttleworth
Honorary Visiting Research Fellow

Affiliations
Links
- https://twitter.com/Welshsquirrels
Personal account - https://twitter.com/RedSquirrelsinW
Welsh red squirrel conservation - https://www.facebook.com/groups/143610382316300
Welsh red squirrel conservation - https://pinemarten.net/
The Gwynedd pine marten project - https://www.linkedin.com/in/craig-shuttleworth-6062a1253/
Personal page
Contact info
Mobile 07966150847
Email c.shuttleworth@bangor.ac.uk
- Published
Why the pine marten is not every red squirrel’s best friend
Shuttleworth, C. & Hayward, M., 2019, The Conversation.Research output: Other contribution
- Published
Wales should follow Scotland’s lead in protecting the red squirrel’s habitat
Shuttleworth, C., 2018, Nation Cymru .Research output: Other contribution
- Published
Virus presence within Welsh red squirrels
Everest, D. J., Holmes, P. & Shuttleworth, C., 30 May 2017, In: The Veterinary record. 180, p. 520-521Research output: Contribution to journal › Article
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Turning the grey tide: progress in red squirrel recovery
Shuttleworth, C., 1 Jun 2010, In: Ecos. 31, p. 27-35Research output: Contribution to journal › Article
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Traffic related mortality in a red squirrel (Sciurus vulgaris) population receiving supplemental feeding
Shuttleworth, C., 1 Jun 2001, In: Urban Ecosystems. 2, p. 109-118Research output: Contribution to journal › Article › peer-review
- Published
The use of nest boxes by the red squirrel Sciurus vulgaris in a coniferous habitat
Shuttleworth, C., 1 Mar 1999, In: Mammal Review. 29, p. 61-66Research output: Contribution to journal › Article › peer-review
- Published
The use of nest boxes by the red squirrel (Sciurus vulgaris) in conifer plantations
Shuttleworth, C., 2001, In: Quarterly Journal of Forestry. 95, p. 225-228Research output: Contribution to journal › Article › peer-review
- Published
The search for novelty continues for rewilding
Hayward, M., Jachowski, J. D., Bugir, C. J., Clulow, J., Krishnamurthy, R., Griffin, A. S., Chalmers, A. C., Linnell, J. D. C., Montgomery, R. A., Somers, M. J., Kowalczyk, R., Heurich, M., Caravaggi, A., Marnewick, K. A., Di Blanco, Y., Shuttleworth, C., Callen, C., Weise, F., Scanlon, R., Moehrenschlager, A., Howell, L. & Upton, R. M. O., 1 Aug 2019, In: Biological Conservation. 236, p. 584-585Research output: Contribution to journal › Article › peer-review
- Published
The role of spruce in red squirrel (Sciurus vulgaris) conservation in the UK
Shuttleworth, C. & Lurz, P. W. W., 2014, Forest Ecosystems: Biodiversity, Management and Conservation. Roberts, N. C. (ed.). Nova Science Publishers, p. 63-78Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review
- Published
The reintroduction of the red squirrel Sciurus vulgaris to Newborough forest, North Wales: A five year project review
Shuttleworth, C., Kenward, R. E. & Jackson, N., 2009Research output: Book/Report › Commissioned report
- Published
The potential role of ectoparasites in the epidemiology of squirrelpox virus: a possible novel means of intervention to reduce the impact of the disease?
Cowan, D. P., Mill, A. C., Everest, D. J., McInnes, C. J., Rushton, S. P., Shirley, M. D. F., Start, C. & Shuttleworth, C., 2016, The grey squirrel: ecology and management in Europe . Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (eds.). European Squirrel Initiative, p. 255-289Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review
- Published
The management of grey squirrel populations in Europe: evolving best practice
Bertolino, S., Lurz, P. W. W., Shuttleworth, C., Martinoli, A. & Wauters, L. A., 2016, The grey squirrel: ecology and management in Europe . Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (eds.). European Squirrel Initiative, p. 495-516Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review
- Published
The management of coastal sand dune woodland for red squirrels (Sciurus vulgaris L.)
Shuttleworth, C. & Gurnell, J., 2001, Coastal Dune Management: Shared Experience of European Conservation Practice. Houston, J., Edmondson, A. S. E. & Rooney, P. J. (eds.). Liverpool University Press, p. 117-127Research output: Chapter in Book/Report/Conference proceeding › Chapter
- Published
The large-scale removal of mammalian invasive alien species in Northern Europe
Robertson, P. A., Adriaens, T., Lambin, X., Mill, A., Roy, S., Shuttleworth, C. & Sutton-Croft, M., 1 Feb 2017, In: Pest Management Science. 73, p. 273-279Research output: Contribution to journal › Article › peer-review
- Published
The implications of significant adenovirus infection in UK captive red squirrel (Sciurus vulgaris) collections: How histological screening can aid applied conservation management
Everest, D. J., Shuttleworth, C., Grierson, S. S., Dastjerdi, A., M.F., S., Duff, J. P., Higgins, R. J., Mill, A. C. & Chantrey, J., 1 Jan 2018, In: Mammalian Biology. 88, p. 123-129Research output: Contribution to journal › Article › peer-review
- Published
The impact of woodland management upon the red squirrel population in Ainsdale National Nature Reserve
Shuttleworth, C., 1997Research output: Book/Report › Commissioned report
- Published
The impact of adenoviruses on red squirrel (Sciurus vulgaris) conservation – with notes on other enteric viruses of potential significance
Everest, D. J., Shuttleworth, C., Grierson, S. S., Dastjerdi, A., Stidworthy, M. F., Duff, J. P. & Sainsbury, A. W., 2015, Red squirrels: Ecology, Conservation & Management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Hayward, M. W. (eds.). European Squirrel Initiative, p. 129-146Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review
- Published
The grey squirrel: ecology & management of an Invasive Species in Europe
Shuttleworth, C., Lurz, P. W. W. & Gurnell, J., 2016, European Squirrel Initiative. 505 p.Research output: Book/Report › Book › peer-review
- Published
The foraging behaviour and diet of red squirrels (Sciurus vulgaris) receiving supplemental feeding
Shuttleworth, C., 1 Sept 2000, In: Wildlife Biology. 6, p. 149-156Research output: Contribution to journal › Article › peer-review
- Published
The effect of supplemental feeding on the diet, population density and reproduction of red squirrels
Shuttleworth, C., 1997, The Conservation of Red Squirrels (Sciurus vulgaris L.). Gurnell, J. & Lurz, P. W. W. (eds.). Peoples Trust for Endangered Species, p. 13-24Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review
- Published
Sympatric cases of squirrelpox in red and grey squirrels
Shuttleworth, C., Everest, D., Wood, C. & Holmes, J. P., 7 May 2021, In: The Veterinary record. 188, 9, p. 358Research output: Contribution to journal › Article › peer-review
- Published
Striving for success: an evaluation of local action to conserve red squirrels (Sciurus vulgaris) in Wales
Halliwell, E. C., Shuttleworth, C., Wilberforce, L., Denman, H., Lloyd, I. & Cartmel, S., 2015, Red squirrels: Ecology, Conservation & Management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Hayward, M. W. (eds.). European Squirrel Initiative, p. 175-192Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review
- Published
Squirrelpox outbreak detected in north Wales – without a vaccine, the disease will keep decimating red squirrels
Shuttleworth, C., 21 Dec 2022, The Conversation.Research output: Other contribution
- Published
Significant differences in the caecal bacterial microbiota of red and grey squirrels in Britain
Hall, L., Nichols, C., Leng, J., Martelli, F., Shuttleworth, C. & La Ragione, R., 14 Feb 2024, In: Journal of Medical Microbiology. 73, 2Research output: Contribution to journal › Article › peer-review
- Published
Shared experience of red squirrel conservation practice
Shuttleworth, C., Lurz, P. W. W. & Halliwell, E. C., 2015, European Squirrel Initiative. 220 p.Research output: Book/Report › Book
- Published
Saving the red squirrel: landscape scale recovery
Shuttleworth, C., Lurz, P. W. W. & Robinson, N., 2021, London: Red Squirrel Survival Trust. 306 p.Research output: Book/Report › Book › peer-review
- Published
Rewilding needs to clarify the role of management of invasive species
Hayward, M., Jachowski, D. S., Shuttleworth, C., Linnell, J. D. C., Allen, B. L., Griffen, A. S., Montgomery, R. A., Caravaggi, A., Weise, F. J., Moehrenschlager, A., Somers, M. J., Clulow, S., Heurich, M. & Marnewick, K., 8 May 2019, In: Science. 364, p. 6438Research output: Contribution to journal › Letter
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Retrospective detection by negative contrast electron microscopy of faecal viral particles in wild red squirrels (Sciurus vulgaris) with suspected enteropathy in Great Britain
Everest, D. J., Stidworthy, M. F., Milne, E. M., Meredith, A. L., Chantery, J., Shuttleworth, C., Blackett, T., Butler, H., Wilkinson, M. & Sainsbury, A. W., 31 Dec 2010, In: Veterinary Record. 167, p. 1007-1010Research output: Contribution to journal › Article › peer-review
- Published
Restoring our lost wildlife: a review of the ecological, social and economic factors in the potential reintroduction of six mammal species to Cornwall
Cooper, P., Brazier, R., Puttock, A., Graham, H., Auster, R., Campbell-Palmer, R., Shuttleworth, C. & Pizzi, R., 2023, 258 p.Research output: Book/Report › Commissioned report
- Published
Response to comments on "Compassionate Conservation deserves a morally serious rather than dismissive response - reply to Callen et al., 2020"
Callen, A., Hayward, M. W., Klop-Toker, K., Allen, B. L., Ballard, G., Beranek, C. T., Broekhuis, F., Bugir, C. K., Clarke, R. H., Clulow, J., Clulow, S., Daltry, J. C., Davies-Mostert, H. T., Di Blanco, Y. E., Dixon, V., Fleming, P. J. S., Howell, L. G., Kerley, G. I. H., Legge, S. M., Lenga, D. J., Major, T., Montgomery, R. A., Moseby, K., Meyer, N., Parker, D. M., Periquet, S., Read, J., Scanlon, R. J., Shuttleworth, C., Tamessar, C. T., Taylor, W. A., Tuft, K., Upton, R. M. O., Valenzuela, M., Witt, R. R. & Wuster, W., 30 Apr 2020, In: Biological Conservation. 244Research output: Contribution to journal › Letter › peer-review
- Published
Repeated squirrelpox infection in Welsh red squirrels
Holmes, J. P., Everest, D. & Shuttleworth, C., 3 Feb 2023, In: Veterinary Record. 192, 3, p. 128-129Research output: Contribution to journal › Letter
- Published
Releasing grey squirrels into the wild
Shuttleworth, C., Bertolino, S., Gill, R., Gurnell, J., Hayward, M., Kenward, R. E., Lawton, C., Lurz, P. W. W., McInnes, C. J., Mill, A., Trotter, S. & Wauters, L. A., 31 Mar 2019, In: Veterinary Record. 184, p. 389-390Research output: Contribution to journal › Article
- Published
Reintroductions and translocations of red squirrels within Europe
Lawton, C., Waters, C. & Shuttleworth, C., 2015, Red squirrels: Ecology, Conservation & Management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Hayward, M. W. (eds.). European Squirrel Initiative, p. 193-210Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review
- Published
Reintroducing rewilding to restoration – Rejecting the search for novelty
Hayward, M. W., Scanlon, R. J., Callen, A., Howell, L. G., Klop-Toker, K. L., Di Blanco, Y., Balkenhol, N., Bugir, C. K., Campbell, L., Caravaggi, A., Chalmers, A. C., Clulow, J., Clulow, S., Cross, P., Gould, J. A., Griffin, A. S., Heurich, M., Howe, B. K., Jachowski, D. S., Jhala, Y. V., Krishnamurthy, R., Kowalczyk, R., Lenga, D. J., Linnell, J. D. C., Marnewick, K. A., Moehrenschlager, A., Montgomery, R. A., Osipova, L., Peneaux, C., Rodger, J. C., Sales, L. P., Seeto, R. G. Y., Shuttleworth, C. M., Somers, M. J., Tamessar, C. T., Upton, R. M. O. & Weise, F. J., 1 May 2019, In: Biological Conservation. 233, p. 255-259 5 p.Research output: Contribution to journal › Article › peer-review
- Published
Reinstating trophic cascades as an applied conservation tool to protect forest ecosystems from invasive grey squirrels (Sciurus carolinensis)
Bamber, J. A., Shuttleworth, C. M., Hayward, M. W. & Everest, D. J., 31 Dec 2020, In: Food Webs. 25, e00164.Research output: Contribution to journal › Article › peer-review
- Published
Reducing negative impacts on non-target species during grey squirrel (Sciurus carolinensis) control activities
Shuttleworth, C., Mill, A. C., Van der Wall, Z., Bertolino, S. & Everest, D. J., 1 Jan 2018, In: Quarterly Journal of Forestry. 112, 1, p. 21-29Research output: Contribution to journal › Article › peer-review
- Published
Red squirrels: ecology, conservation & management in Europe
Shuttleworth, C., Hayward, M. & Lurz, P. W. W., 2015, European Squirrel Initiative. 330 p.Research output: Book/Report › Book › peer-review
- Published
Red squirrels on Anglesey
Shuttleworth, C. & Bailey, M., 2003, In: Natur Cymru. 6, p. 35-37Research output: Contribution to journal › Article
- Published
Red squirrels in my garden: guidance and tips to help conserve local populations
Shuttleworth, C. & Halliwell, E. C., 2018, European Squirrel Initiative. 185 p.Research output: Book/Report › Book › peer-review
- Published
Red squirrels (Sciurus vulgaris) on the Sefton Coast: The conservation value of nest boxes
Shuttleworth, C., 1995, 2nd NPI Red Alert UK Forum for Red Squirrel Conservation. Hughes, D. G. & Tew, T. (eds.). The Federation of Zoological Gardens of Great Britain and Ireland, p. 25-32Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review
- Published
Red squirrel population monitoring: Clocaenog forest
Shuttleworth, C. & Donoyou, T., 2012, IFCW108/001/002 ed.Research output: Book/Report › Commissioned report
- Published
Red squirrel conservation on Anglesey, North Wales. A report on the first five years of the Anglesey Red Squirrel Project
Shuttleworth, C., Bailey, M. & Knott, H., 2002Research output: Book/Report › Commissioned report
- Published
Red squirrel conservation and disease - Adenovirus in Wales
Shuttleworth, C., 2014, In: Natur Cymru. 52, p. 41-44Research output: Contribution to journal › Article
- Published
Red squirrel bark-stripping of hornbeam in North Wales
Shuttleworth, C. & Gill, R., 2019, In: Quarterly Journal of Forestry. 113, 4, p. 275-277Research output: Contribution to journal › Article › peer-review
- Published
Red squirrel (Sciurus vulgaris) conservation plan for Anglesey & Gwynedd
Shuttleworth, C. & Halliwell, E. C., 2018, 50 p.Research output: Book/Report › Commissioned report
- Published
Red Squirrel Conservation in Clocaenog Forest: Review and Recommendations
Shuttleworth, C., 2021, 56 p.Research output: Book/Report › Commissioned report
- Published
Recalibrating Risk: Implications of squirrelpox virus for successful red squirrel translocations within mainland UK
Shuttleworth, C., Brady, D., Cross, P., Gardner, L., Greenwood, A., Jackson, N., McKinney, C., Robinson, N., Trotter, S., Valle, S., Wood, K. & Hayward, M., Feb 2021, In: Conservation Science and Practice. 3, 2, e321.Research output: Contribution to journal › Article › peer-review
- Published
Re-introduction of the red squirrel into Newborough forest on the island of Anglesey, UK
Shuttleworth, C., Kenward, R. E. & Jackson, N., 2008, Global Re-Introduction Perspectives: Re-Introduction Case-Studies from Around the Globe. Soorae, P. S. (ed.). IUCN/SSC Re-introduction Specialist Group, p. 163-166Research output: Chapter in Book/Report/Conference proceeding › Chapter
- Published
Rare woodland wildlife at risk because of 50-year-old tree felling rules
Shuttleworth, C., 2018, The Conversation.Research output: Other contribution
- Published
Ranging behaviour, density and social structure in grey squirrels
Lawton, C., Shuttleworth, C. & Kenward, R. E., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (eds.). European Squirrel Initiative, p. 133-152Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review