Dr Craig Shuttleworth

Honorary Visiting Research Fellow

Contact info

Mobile 07966150847

Email c.shuttleworth@bangor.ac.uk

  1. Letter › Research › Peer-reviewed
  2. Published

    Response to comments on "Compassionate Conservation deserves a morally serious rather than dismissive response - reply to Callen et al., 2020"

    Callen, A., Hayward, M. W., Klop-Toker, K., Allen, B. L., Ballard, G., Beranek, C. T., Broekhuis, F., Bugir, C. K., Clarke, R. H., Clulow, J., Clulow, S., Daltry, J. C., Davies-Mostert, H. T., Di Blanco, Y. E., Dixon, V., Fleming, P. J. S., Howell, L. G., Kerley, G. I. H., Legge, S. M., Lenga, D. J., Major, T., Montgomery, R. A., Moseby, K., Meyer, N., Parker, D. M., Periquet, S., Read, J., Scanlon, R. J., Shuttleworth, C., Tamessar, C. T., Taylor, W. A., Tuft, K., Upton, R. M. O., Valenzuela, M., Witt, R. R. & Wuster, W., 30 Apr 2020, In: Biological Conservation. 244

    Research output: Contribution to journalLetterpeer-review

  3. Letter › Research › Not peer-reviewed
  4. Published

    Adenovirus remains widespread in squirrels on Anglesey

    Shuttleworth, C., Everest, D. & McQueen, A., 16 Dec 2023, In: Veterinary Record. 193, 12, p. 490-491 2 p.

    Research output: Contribution to journalLetter

  5. Published

    By trying to be everything, rewilding risks meaning nothing

    Jachowski, J. D., Bombaci, S., Caravaggi, A., Chalmers, A., Clulow, S., Griffin, A., Hayward, M., Heurich, M., Linnell, J., Marnewick, K., Moehrenschlager, A., Montgomery, A., Shuttleworth, C., Somers, M. & Weise, F., Nov 2019, In: Science. 364, p. 6438

    Research output: Contribution to journalLetter

  6. Published

    Repeated squirrelpox infection in Welsh red squirrels

    Holmes, J. P., Everest, D. & Shuttleworth, C., 3 Feb 2023, In: Veterinary Record. 192, 3, p. 128-129

    Research output: Contribution to journalLetter

  7. Published

    Rewilding needs to clarify the role of management of invasive species

    Hayward, M., Jachowski, D. S., Shuttleworth, C., Linnell, J. D. C., Allen, B. L., Griffen, A. S., Montgomery, R. A., Caravaggi, A., Weise, F. J., Moehrenschlager, A., Somers, M. J., Clulow, S., Heurich, M. & Marnewick, K., 8 May 2019, In: Science. 364, p. 6438

    Research output: Contribution to journalLetter

  8. Article › Research › Peer-reviewed
  9. Published

    A preliminary comparison of magnum 116 body-grip traps with a live capture trap design during the control of grey squirrels (Sciurus carolinensis)

    Shuttleworth, C., Mill, A. & Van der Waal, Z., Dec 2016, In: International Journal of Pest Management. 63, 4, p. 316-321

    Research output: Contribution to journalArticlepeer-review

  10. Published

    A study of nest box use by a low density red squirrel (Sciurus vulgaris) population within a commercial conifer plantation

    Shuttleworth, C., 2004, In: Quarterly Journal of Forestry. 98, p. 113-119

    Research output: Contribution to journalArticlepeer-review

  11. Published

    A systematic assessment of the impact of adenovirus infection on a captive re-introduction project for red squirrels (Sciurus vulgaris)

    Everest, D. J., Shuttleworth, C., Grierson, S. S., Duff, J. P., Jackson, N., Litherland, P. & Kenward, R. E., 31 Aug 2012, In: Veterinary Record. 171, p. 176-184

    Research output: Contribution to journalArticlepeer-review

  12. Published

    A temporal refuge from predation can change the outcome of prey species competition

    Slade, A., White, A., Lurz, P. W. W., Shuttleworth, C. & Lambin, X., 1 Sept 2022, In: Oikos. Oikos e08565

    Research output: Contribution to journalArticlepeer-review

  13. Published

    A tough nut to crack: red squirrel conservation in Wales

    Shuttleworth, C., 1 Oct 2003, In: Biologist. 50, p. 231-235

    Research output: Contribution to journalArticlepeer-review

  14. Published

    Adenovirus infection: An emerging factor in red squirrel conservation programmes

    Everest, D. J., Shuttleworth, C., Stidworthy, M. F., Grierson, S. S., Duff, P. J. & Kenward, R. E., 1 Oct 2014, In: Mammal Review. 44, 3-4, p. 225-233

    Research output: Contribution to journalArticlepeer-review

  15. Published

    Adenovirus-associated deaths in red squirrels on Anglesey

    Everest, D. J., Shuttleworth, C. & Stidworthy, M. F., 31 Oct 2008, In: The Veterinary record. 163, 14, p. 430

    Research output: Contribution to journalArticlepeer-review

  16. Published

    Against a grey tide: red squirrel conservation on Anglesey

    Shuttleworth, C., 2005, In: Quarterly Journal of Forestry. 99, p. 213-214

    Research output: Contribution to journalArticlepeer-review

  17. Published

    An Opportunistic Assessment of the Impact of Squirrelpox Disease Outbreaks upon a Red Squirrel Population Sympatric with Grey Squirrels in Wales

    Shuttleworth, C., Everest, D., Holmes, P., Bell, S. & Cripps, R., 1 Jan 2022, In: Animals. 12, 1

    Research output: Contribution to journalArticlepeer-review

  18. Published

    Are nest boxes a useful tool in regional red squirrel conservation programs?

    Shuttleworth, C. & Schuchert, P., 15 Dec 2014, In: Hystrix-Italian Journal of Mammalogy. 25, p. 91-94

    Research output: Contribution to journalArticlepeer-review

  19. Published

    Are novel ecosystems the only novelty of rewilding?

    Klop-Toker, K., Clulow, S., Shuttleworth, C. & Hayward, M., 1 Nov 2020, In: Restoration Ecology. 28, p. 1318-1320

    Research output: Contribution to journalArticlepeer-review

  20. Published

    Assessing a potential non-invasive method for viral diagnostic purposes in European squirrels

    Everest, D. J., Tolhurst-Cherriman, D. A. R., Davies, H., Dastjerdi, A., Ashton, A., Blackett, T., Meredith, A. L., Milne, E., Mill, A. & Shuttleworth, C., 4 Jun 2019, In: Hystrix-Italian Journal of Mammalogy. 30, p. 44-50

    Research output: Contribution to journalArticlepeer-review

  21. Published

    Assessing causes and significance of red squirrel (Sciurus vulgaris) mortality during regional population restoration: An applied conservation perspective

    Shuttleworth, C., Signorile, A. L., Everest, D. J., Lurz, P. W. W. & Duff, J. P., 16 Nov 2015, In: Hystrix-Italian Journal of Mammalogy. 26, 2, p. 69-75

    Research output: Contribution to journalArticlepeer-review

  22. Published

    British red squirrels remain the only known wild rodent host for leprosy bacilli

    Schilling, A., Avanzi, C., Ulrich, R. G., Busso, P., Pisanu, B., Ferrari, N., Romeo, C., Mazzamuto, M. V., McLuckie, J., Shuttleworth, C., Del-Pozo, J., Lurz, P. W. W., Escalante-Fuentes, W. G., Ocampo-Candiani, J., Vera-Cabrera, L., Stevenson, K., Chapuis, J-L., Meredith, A. L. & Cole, S. T., 1 Feb 2019, In: Frontiers in Veterinary Science. 6, p. 8

    Research output: Contribution to journalArticlepeer-review

  23. Published

    Capturing red squirrels (Sciurus vulgaris) on camera: A cost-effective approach for monitoring relative abundance and habitat preference

    Shannon, G., Valle, S. & Shuttleworth, C., 3 Oct 2023, In: Ecology and Evolution. 13, 10, e10536.

    Research output: Contribution to journalArticlepeer-review

  24. Published

    Challenges and solutions in early detection, rapid response and communication about potential invasive alien species in forests

    de Groot, M., Bullas-Appleton, E., Csóka, G., Csiszár, A., Faccoli, A., Gervasini, A., Kirichenko, N., Shuttleworth, C., Kus Veenvliet, J., O'Hanlon, R., Korda, M., Marinsek, A., Robinson, N., Sweeney, J., Tricarico, E., Verbrugge, L., Williams, D. & Zidar, S., 14 Aug 2020, In: Management of Biological Invasions. 11, p. 637-660

    Research output: Contribution to journalArticlepeer-review

  25. Published

    Deconstructing compassionate conservation

    Hayward, M. W., Callen, A., Allen, B. L., Ballard, G., Broekhuis, F., Bugir, C., Clarke, R. H., Clulow, J., Clulow, S., Daltry, J. C., Davies-Mostert, H. T., Fleming, P. J. S., Griffin, A. S., Howell, L. G., Kerley, G. I. H., Klop-Toker, K., Legge, S., Major, T., Meyer, N., Montgomery, R. A., Moseby, K., Parker, D. M., Périquet, S., Read, J., Scanlon, R., Seeto, R., Shuttleworth, C., Somers, M. J., Tamessar, C. T., Tuft, K., Upton, R., Valenzuela-Molina, M., Wayne, A., Witt, R. R. & Wuster, W., Aug 2019, In: Conservation Biology. 33, 4, p. 760-768

    Research output: Contribution to journalArticlepeer-review

  26. Published

    Disease monitoring and surveillance: case studies in the applied conservation of fragmented red squirrel (Sciurus vulgaris) populations in England and Wales

    Everest, D. J., Floyd, T., Holmes, P., Man, C., Dunnett, C., Locke, R., Savage, L., Sutcliffe, S., Sapsford, B., Shuttleworth, C. & Duff, P., 1 Dec 2021, In: Mammalian Biology. 101, p. 1079-1088

    Research output: Contribution to journalArticlepeer-review

  27. Published

    Disease, invasions and conservation: no evidence of squirrelpox virus in grey squirrels introduced to Italy

    Romeo, C., McInnes, C., Dale, T., Shuttleworth, C., Bertolino, S., Wauters, L. A. & Ferrari, N., 1 Feb 2019, In: Animal Conservation. 22, 1, p. 14-23

    Research output: Contribution to journalArticlepeer-review

  28. Published

    Do differing levels of boldness influence the success of translocation? A pilot study on red squirrels (Sciurus vulgaris)

    Bamber, J. A., Shuttleworth, C. & Hayward, M., 26 Sept 2020, In: Animals. 10, 1748.

    Research output: Contribution to journalArticlepeer-review

  29. Published

    Envisioning the future with 'compassionate conservation': An ominous projection for native wildlife and biodiversity

    Callen, A., Hayward, M. W., Klop-Toker, K., Allen, B. L., Ballard, G., Beranek, C. T., Broekhuis, F., Bugir, C. K., Clarke, R. H., Clulow, J., Clulow, S., Daltry, J. C., Davies-Mostert, H. T., Di Blanco, Y. E., Dixon, V., Fleming, P. J. S., Howell, L. G., Kerley, G. I. H., Legge, S. M., Lenga, D. J., Major, T., Montgomery, R. A., Moseby, K., Meyer, N., Parker, D. M., Periquet, S., Read, J., Scanlon, R. J., Shuttleworth, C., Tamessar, C. T., Taylor, W. A., Tuft, K., Upton, R. M. O., Valenzuela, M., Witt, R. R. & Wuster, W., Jan 2020, In: Biological Conservation. 241, 108365.

    Research output: Contribution to journalArticlepeer-review

  30. Published

    Ethical dilemmas when using citizen science for early detection of invasive tree pests and diseases

    Pocock, M. J. O., Marzano, M., Bullas-Appleton, E., Dyke, A., de Groot, M., Shuttleworth, C. & White, R., 1 Nov 2020, In: Management of Biological Invasions. 11, p. 720-732

    Research output: Contribution to journalArticlepeer-review

  31. Published

    Evolving grey squirrel management techniques in Europe

    Shuttleworth, C., Robinson, N. J., Halliwell, E. C., Clews-Roberts, R., Peek, H., Podgornik, G., Stinson, M., Rice, S., Finlay, C., McKinney, C., Everest, D. J. & Larsen, K. W., 21 Oct 2020, In: Management of Biological Invasions. 11, 4, p. 747-761

    Research output: Contribution to journalArticlepeer-review

  32. Published

    First report of dermatophilosis in wild European red squirrels (Sciurus vulgaris)

    Holmes, P., Everest, D. J., Spiro, S., Wessels, M. & Shuttleworth, C., 30 Sept 2019, In: Veterinary Record Case Reports. 7, 3, p. 3

    Research output: Contribution to journalArticlepeer-review

  33. Published

    Genetic management of the red squirrel, Sciurus vulgaris: a practical approach to regional conservation

    Ogden, R., Shuttleworth, C., McEwing, R. & Cesarini, S., 26 Oct 2005, In: Conservation Genetics. 6, p. 511-525

    Research output: Contribution to journalArticlepeer-review

  34. Published

    Indirect effects of pine marten recovery result in benefits to native prey through suppression of an invasive species and a shared pathogen

    Slade, A., White, A., Lurz, P. W. W., Shuttleworth, C., Tosh, D. & Twining, JP., 1 Feb 2023, In: Ecological Modelling. 476, 110216.

    Research output: Contribution to journalArticlepeer-review

  35. Published

    Integrating red squirrel (Sciurus vulgaris) habitat requirements with the management of pathogenic tree disease in commercial forests in the UK

    Shuttleworth, C., Lurz, P. W. W., Geddes, N. & Brown, J., 1 Sept 2012, In: Forest Ecology and Management. 279, p. 167-175

    Research output: Contribution to journalArticlepeer-review

  36. Published

    Inter-specific viral infections: Can the management of captive red squirrel collections help inform scientific research?

    Shuttleworth, C., Everest, D. J., McInnes, C. J., Greenwood, A., Jackson, N., Rushton, S. P. & Kenward, R. E., 16 Jun 2014, In: Hystrix-Italian Journal of Mammalogy. 25, 1, p. 18-24

    Research output: Contribution to journalArticlepeer-review

  37. Published

    Interactions between the red squirrel (Sciurus vulgaris), great tit (Parus Major) and jackdaw (Corvus monedula) whilst using nest boxes.

    Shuttleworth, C., 1 Oct 2001, In: Journal of Zoology. 255, p. 269-272

    Research output: Contribution to journalArticlepeer-review

  38. Published

    Introduced Canadian Eastern grey squirrels: squirrelpox virus surveillance and why nothing matters.

    McInnes, C., Shuttleworth, C., Larsen, K. W., Everest, D. J., Bruemmer, C., Carroll, B., Romeo, C., Sainsbury, A. W., Crawshaw, G., Dubois, S., Gillis, L., Gilray, J. & Percival, A., 3 Aug 2020, In: Hystrix-Italian Journal of Mammalogy. 31, p. 95-98

    Research output: Contribution to journalArticlepeer-review

  39. E-pub ahead of print

    Invasive genetic rescue: dispersal following repeated culling reinforces the genetic diversity of an invasive mammal

    Synnott, R., Shuttleworth, C., Everest, D., Stevenson-Holt, C., O'Reilly, C., McDevitt, A. & O.Meara, D., 30 Jun 2023, (E-pub ahead of print) In: Biological Invasions.

    Research output: Contribution to journalArticlepeer-review

  40. Published

    Landscape scale impacts of culling upon a European grey squirrel population: can trapping reduce population size and decrease the threat of squirrel pox virus infection for the native red squirrel?

    Schuchert, P., Shuttleworth, C., McInnes, C. J., Everest, D. J. & Rushton, S. P., 1 Nov 2014, In: Biological Invasions. 16, p. 2381-2391

    Research output: Contribution to journalArticlepeer-review

  41. Published

    Mathematical models for invasive species management: grey squirrel control on Anglesey

    Jones, H., White, A., Lurz, P. W. W. & Shuttleworth, C., 10 Sept 2017, In: Ecological Modelling. 180, p. 274-284

    Research output: Contribution to journalArticlepeer-review

  42. Published

    Opportunistic viral surveillance confirms the ongoing disease threat grey squirrels pose to sympatric red squirrel populations in the UK

    Everest, DJ., Green, C., Dastjerdi, A., Davies, H., Cripps, R., McKinney, C., Podgornik, G., Stinson, M., O'Hare, S., Bonnie, S., Mill, A., Van der wall, Z., Robinson, N., Trotter, S. & Shuttleworth, C., 20 May 2023, In: Veterinary Record. 192, 10, e2834.

    Research output: Contribution to journalArticlepeer-review

  43. Published

    PCR detection of adenovirus in grey squirrels on Anglesey

    Everest, D. J., Grierson, S. S., Stidworthy, M. & Shuttleworth, C., 31 Oct 2009, In: Veterinary Record. 165, p. 482

    Research output: Contribution to journalArticlepeer-review

  44. Published

    Recalibrating Risk: Implications of squirrelpox virus for successful red squirrel translocations within mainland UK

    Shuttleworth, C., Brady, D., Cross, P., Gardner, L., Greenwood, A., Jackson, N., McKinney, C., Robinson, N., Trotter, S., Valle, S., Wood, K. & Hayward, M., Feb 2021, In: Conservation Science and Practice. 3, 2, e321.

    Research output: Contribution to journalArticlepeer-review

  45. Published

    Red squirrel bark-stripping of hornbeam in North Wales

    Shuttleworth, C. & Gill, R., 2019, In: Quarterly Journal of Forestry. 113, 4, p. 275-277

    Research output: Contribution to journalArticlepeer-review

  46. Published

    Reducing negative impacts on non-target species during grey squirrel (Sciurus carolinensis) control activities

    Shuttleworth, C., Mill, A. C., Van der Wall, Z., Bertolino, S. & Everest, D. J., 1 Jan 2018, In: Quarterly Journal of Forestry. 112, 1, p. 21-29

    Research output: Contribution to journalArticlepeer-review

  47. Published

    Reinstating trophic cascades as an applied conservation tool to protect forest ecosystems from invasive grey squirrels (Sciurus carolinensis)

    Bamber, J. A., Shuttleworth, C. M., Hayward, M. W. & Everest, D. J., 31 Dec 2020, In: Food Webs. 25, e00164.

    Research output: Contribution to journalArticlepeer-review

  48. Published

    Reintroducing rewilding to restoration – Rejecting the search for novelty

    Hayward, M. W., Scanlon, R. J., Callen, A., Howell, L. G., Klop-Toker, K. L., Di Blanco, Y., Balkenhol, N., Bugir, C. K., Campbell, L., Caravaggi, A., Chalmers, A. C., Clulow, J., Clulow, S., Cross, P., Gould, J. A., Griffin, A. S., Heurich, M., Howe, B. K., Jachowski, D. S., Jhala, Y. V., Krishnamurthy, R., Kowalczyk, R., Lenga, D. J., Linnell, J. D. C., Marnewick, K. A., Moehrenschlager, A., Montgomery, R. A., Osipova, L., Peneaux, C., Rodger, J. C., Sales, L. P., Seeto, R. G. Y., Shuttleworth, C. M., Somers, M. J., Tamessar, C. T., Upton, R. M. O. & Weise, F. J., 1 May 2019, In: Biological Conservation. 233, p. 255-259 5 p.

    Research output: Contribution to journalArticlepeer-review

  49. Retrospective detection by negative contrast electron microscopy of faecal viral particles in wild red squirrels (Sciurus vulgaris) with suspected enteropathy in Great Britain

    Everest, D. J., Stidworthy, M. F., Milne, E. M., Meredith, A. L., Chantery, J., Shuttleworth, C., Blackett, T., Butler, H., Wilkinson, M. & Sainsbury, A. W., 31 Dec 2010, In: Veterinary Record. 167, p. 1007-1010

    Research output: Contribution to journalArticlepeer-review

  50. Published

    Significant differences in the caecal bacterial microbiota of red and grey squirrels in Britain

    Hall, L., Nichols, C., Leng, J., Martelli, F., Shuttleworth, C. & La Ragione, R., 14 Feb 2024, In: Journal of Medical Microbiology. 73, 2

    Research output: Contribution to journalArticlepeer-review

  51. Published

    Sympatric cases of squirrelpox in red and grey squirrels

    Shuttleworth, C., Everest, D., Wood, C. & Holmes, J. P., 7 May 2021, In: The Veterinary record. 188, 9, p. 358

    Research output: Contribution to journalArticlepeer-review

  52. Published

    The foraging behaviour and diet of red squirrels (Sciurus vulgaris) receiving supplemental feeding

    Shuttleworth, C., 1 Sept 2000, In: Wildlife Biology. 6, p. 149-156

    Research output: Contribution to journalArticlepeer-review

  53. Published

    The implications of significant adenovirus infection in UK captive red squirrel (Sciurus vulgaris) collections: How histological screening can aid applied conservation management

    Everest, D. J., Shuttleworth, C., Grierson, S. S., Dastjerdi, A., M.F., S., Duff, J. P., Higgins, R. J., Mill, A. C. & Chantrey, J., 1 Jan 2018, In: Mammalian Biology. 88, p. 123-129

    Research output: Contribution to journalArticlepeer-review