Dr Craig Shuttleworth

Honorary Visiting Research Fellow

Contact info

Mobile 07966150847

Email c.shuttleworth@bangor.ac.uk

  1. Published

    Do differing levels of boldness influence the success of translocation? A pilot study on red squirrels (Sciurus vulgaris)

    Bamber, J. A., Shuttleworth, C. & Hayward, M., 26 Sept 2020, In: Animals. 10, 1748.

    Research output: Contribution to journalArticlepeer-review

  2. Published

    Reinstating trophic cascades as an applied conservation tool to protect forest ecosystems from invasive grey squirrels (Sciurus carolinensis)

    Bamber, J. A., Shuttleworth, C. M., Hayward, M. W. & Everest, D. J., 31 Dec 2020, In: Food Webs. 25, e00164.

    Research output: Contribution to journalArticlepeer-review

  3. Published

    The management of grey squirrel populations in Europe: evolving best practice

    Bertolino, S., Lurz, P. W. W., Shuttleworth, C., Martinoli, A. & Wauters, L. A., 2016, The grey squirrel: ecology and management in Europe . Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (eds.). European Squirrel Initiative, p. 495-516

    Research output: Chapter in Book/Report/Conference proceedingChapterpeer-review

  4. Published

    Response to comments on "Compassionate Conservation deserves a morally serious rather than dismissive response - reply to Callen et al., 2020"

    Callen, A., Hayward, M. W., Klop-Toker, K., Allen, B. L., Ballard, G., Beranek, C. T., Broekhuis, F., Bugir, C. K., Clarke, R. H., Clulow, J., Clulow, S., Daltry, J. C., Davies-Mostert, H. T., Di Blanco, Y. E., Dixon, V., Fleming, P. J. S., Howell, L. G., Kerley, G. I. H., Legge, S. M., Lenga, D. J., Major, T., Montgomery, R. A., Moseby, K., Meyer, N., Parker, D. M., Periquet, S., Read, J., Scanlon, R. J., Shuttleworth, C., Tamessar, C. T., Taylor, W. A., Tuft, K., Upton, R. M. O., Valenzuela, M., Witt, R. R. & Wuster, W., 30 Apr 2020, In: Biological Conservation. 244

    Research output: Contribution to journalLetterpeer-review

  5. Published

    Envisioning the future with 'compassionate conservation': An ominous projection for native wildlife and biodiversity

    Callen, A., Hayward, M. W., Klop-Toker, K., Allen, B. L., Ballard, G., Beranek, C. T., Broekhuis, F., Bugir, C. K., Clarke, R. H., Clulow, J., Clulow, S., Daltry, J. C., Davies-Mostert, H. T., Di Blanco, Y. E., Dixon, V., Fleming, P. J. S., Howell, L. G., Kerley, G. I. H., Legge, S. M., Lenga, D. J., Major, T., Montgomery, R. A., Moseby, K., Meyer, N., Parker, D. M., Periquet, S., Read, J., Scanlon, R. J., Shuttleworth, C., Tamessar, C. T., Taylor, W. A., Tuft, K., Upton, R. M. O., Valenzuela, M., Witt, R. R. & Wuster, W., Jan 2020, In: Biological Conservation. 241, 108365.

    Research output: Contribution to journalArticlepeer-review

  6. Published

    Restoring our lost wildlife: a review of the ecological, social and economic factors in the potential reintroduction of six mammal species to Cornwall

    Cooper, P., Brazier, R., Puttock, A., Graham, H., Auster, R., Campbell-Palmer, R., Shuttleworth, C. & Pizzi, R., 2023, 258 p.

    Research output: Book/ReportCommissioned report

  7. Published

    The potential role of ectoparasites in the epidemiology of squirrelpox virus: a possible novel means of intervention to reduce the impact of the disease?

    Cowan, D. P., Mill, A. C., Everest, D. J., McInnes, C. J., Rushton, S. P., Shirley, M. D. F., Start, C. & Shuttleworth, C., 2016, The grey squirrel: ecology and management in Europe . Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (eds.). European Squirrel Initiative, p. 255-289

    Research output: Chapter in Book/Report/Conference proceedingChapterpeer-review

  8. Published

    Non-invasive genetic studies and virus epidemiology

    Everest, D. J., Dastjerdi, A., Davies, H., Shuttleworth, C., Ogden, R., Butler, H. & Sapsford, B., 30 Mar 2022, In: Veterinary Record. 190, p. 204-205

    Research output: Contribution to journalArticle

  9. Published

    Adenovirus-associated deaths in red squirrels on Anglesey

    Everest, D. J., Shuttleworth, C. & Stidworthy, M. F., 31 Oct 2008, In: The Veterinary record. 163, 14, p. 430

    Research output: Contribution to journalArticlepeer-review

  10. Published

    Virus presence within Welsh red squirrels

    Everest, D. J., Holmes, P. & Shuttleworth, C., 30 May 2017, In: The Veterinary record. 180, p. 520-521

    Research output: Contribution to journalArticle

  11. Published

    The implications of significant adenovirus infection in UK captive red squirrel (Sciurus vulgaris) collections: How histological screening can aid applied conservation management

    Everest, D. J., Shuttleworth, C., Grierson, S. S., Dastjerdi, A., M.F., S., Duff, J. P., Higgins, R. J., Mill, A. C. & Chantrey, J., 1 Jan 2018, In: Mammalian Biology. 88, p. 123-129

    Research output: Contribution to journalArticlepeer-review

  12. Published

    Opportunistic viral surveillance confirms the ongoing disease threat grey squirrels pose to sympatric red squirrel populations in the UK

    Everest, DJ., Green, C., Dastjerdi, A., Davies, H., Cripps, R., McKinney, C., Podgornik, G., Stinson, M., O'Hare, S., Bonnie, S., Mill, A., Van der wall, Z., Robinson, N., Trotter, S. & Shuttleworth, C., 20 May 2023, In: Veterinary Record. 192, 10, e2834.

    Research output: Contribution to journalArticlepeer-review

  13. Published

    Assessing a potential non-invasive method for viral diagnostic purposes in European squirrels

    Everest, D. J., Tolhurst-Cherriman, D. A. R., Davies, H., Dastjerdi, A., Ashton, A., Blackett, T., Meredith, A. L., Milne, E., Mill, A. & Shuttleworth, C., 4 Jun 2019, In: Hystrix-Italian Journal of Mammalogy. 30, p. 44-50

    Research output: Contribution to journalArticlepeer-review

  14. Published

    The impact of adenoviruses on red squirrel (Sciurus vulgaris) conservation – with notes on other enteric viruses of potential significance

    Everest, D. J., Shuttleworth, C., Grierson, S. S., Dastjerdi, A., Stidworthy, M. F., Duff, J. P. & Sainsbury, A. W., 2015, Red squirrels: Ecology, Conservation & Management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Hayward, M. W. (eds.). European Squirrel Initiative, p. 129-146

    Research output: Chapter in Book/Report/Conference proceedingChapterpeer-review

  15. Published

    Adenovirus infection: An emerging factor in red squirrel conservation programmes

    Everest, D. J., Shuttleworth, C., Stidworthy, M. F., Grierson, S. S., Duff, P. J. & Kenward, R. E., 1 Oct 2014, In: Mammal Review. 44, 3-4, p. 225-233

    Research output: Contribution to journalArticlepeer-review

  16. Retrospective detection by negative contrast electron microscopy of faecal viral particles in wild red squirrels (Sciurus vulgaris) with suspected enteropathy in Great Britain

    Everest, D. J., Stidworthy, M. F., Milne, E. M., Meredith, A. L., Chantery, J., Shuttleworth, C., Blackett, T., Butler, H., Wilkinson, M. & Sainsbury, A. W., 31 Dec 2010, In: Veterinary Record. 167, p. 1007-1010

    Research output: Contribution to journalArticlepeer-review

  17. Published

    Confirmation of squirrelpox in Welsh red squirrel

    Everest, D. J., Floyd, T., Donnnacie, B., Irvine, R. M., Holmes, J. P. & Shuttleworth, C., 30 Nov 2017, In: Veterinary Record. 181, p. 514-515

    Research output: Contribution to journalArticle

  18. Published

    A systematic assessment of the impact of adenovirus infection on a captive re-introduction project for red squirrels (Sciurus vulgaris)

    Everest, D. J., Shuttleworth, C., Grierson, S. S., Duff, J. P., Jackson, N., Litherland, P. & Kenward, R. E., 31 Aug 2012, In: Veterinary Record. 171, p. 176-184

    Research output: Contribution to journalArticlepeer-review

  19. Published

    Disease monitoring and surveillance: case studies in the applied conservation of fragmented red squirrel (Sciurus vulgaris) populations in England and Wales

    Everest, D. J., Floyd, T., Holmes, P., Man, C., Dunnett, C., Locke, R., Savage, L., Sutcliffe, S., Sapsford, B., Shuttleworth, C. & Duff, P., 1 Dec 2021, In: Mammalian Biology. 101, p. 1079-1088

    Research output: Contribution to journalArticlepeer-review

  20. Published

    PCR detection of adenovirus in grey squirrels on Anglesey

    Everest, D. J., Grierson, S. S., Stidworthy, M. & Shuttleworth, C., 31 Oct 2009, In: Veterinary Record. 165, p. 482

    Research output: Contribution to journalArticlepeer-review

  21. Published

    Adenovirus detections in juvenile squirrels

    Everest, D. J., Dastjerdi, A., Cowan, D., Gomm, M., Shuttleworth, C., McInnes, C. J., Deane, D., Coulter, L., Mill, A., Rushton, S. P., Jackson, N. L. & Litherland, P., 1 Oct 2015, In: Veterinary Record. 177, p. 373-374

    Research output: Contribution to journalArticle

  22. Published

    Adenovirus infection in red squirrels in areas free from grey squirrels

    Everest, D. J., Butler, H., Simpson, V. R., Blackett, T. & Shuttleworth, C., 31 Aug 2013, In: The Veterinary record. 173, 8, p. 199-200

    Research output: Contribution to journalArticle

  23. Published

    Controlling grey squirrels in forests and woodlands in the UK

    Gill, R., Ferryman, M., Shuttleworth, C., Lurz, P. W. W., Mill, A., Robertson, P. A. & Dutton, C., 2019, Forest Research. 16 p.

    Research output: Book/ReportBookpeer-review

  24. Published

    British red squirrel strongholds: challenges for conservation

    Gurnell, J., Blackett, T., Butler, H., Lurz, P. W. W., Magris, L. & Shuttleworth, C., 2015, Red squirrels: Ecology, Conservation & Management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Hayward, M. W. (eds.). European Squirrel Initiative, p. 211-232

    Research output: Chapter in Book/Report/Conference proceedingChapterpeer-review

  25. Published

    Ecosystem impacts of an alien invader in Europe, the grey squirrel Sciurus carolinensis

    Gurnell, J., Lurz, P. W. W. & Shuttleworth, C., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (eds.). European Squirrel Initiative, p. 307-329

    Research output: Chapter in Book/Report/Conference proceedingChapterpeer-review

  26. Published

    Significant differences in the caecal bacterial microbiota of red and grey squirrels in Britain

    Hall, L., Nichols, C., Leng, J., Martelli, F., Shuttleworth, C. & La Ragione, R., 14 Feb 2024, In: Journal of Medical Microbiology. 73, 2

    Research output: Contribution to journalArticlepeer-review

  27. Published

    Striving for success: an evaluation of local action to conserve red squirrels (Sciurus vulgaris) in Wales

    Halliwell, E. C., Shuttleworth, C., Wilberforce, L., Denman, H., Lloyd, I. & Cartmel, S., 2015, Red squirrels: Ecology, Conservation & Management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Hayward, M. W. (eds.). European Squirrel Initiative, p. 175-192

    Research output: Chapter in Book/Report/Conference proceedingChapterpeer-review

  28. Published

    Experiences of grey squirrel management in an upland conifer forest

    Halliwell, E. C., Shuttleworth, C., Cartmel, S., Lloyd, I. & Jenkins, R., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (eds.). European Squirrel Initiative, p. 453-474

    Research output: Chapter in Book/Report/Conference proceedingChapterpeer-review

  29. Published

    Rewilding needs to clarify the role of management of invasive species

    Hayward, M., Jachowski, D. S., Shuttleworth, C., Linnell, J. D. C., Allen, B. L., Griffen, A. S., Montgomery, R. A., Caravaggi, A., Weise, F. J., Moehrenschlager, A., Somers, M. J., Clulow, S., Heurich, M. & Marnewick, K., 8 May 2019, In: Science. 364, p. 6438

    Research output: Contribution to journalLetter

  30. Published

    The search for novelty continues for rewilding

    Hayward, M., Jachowski, J. D., Bugir, C. J., Clulow, J., Krishnamurthy, R., Griffin, A. S., Chalmers, A. C., Linnell, J. D. C., Montgomery, R. A., Somers, M. J., Kowalczyk, R., Heurich, M., Caravaggi, A., Marnewick, K. A., Di Blanco, Y., Shuttleworth, C., Callen, C., Weise, F., Scanlon, R., Moehrenschlager, A., Howell, L. & Upton, R. M. O., 1 Aug 2019, In: Biological Conservation. 236, p. 584-585

    Research output: Contribution to journalArticlepeer-review

  31. Published

    Reintroducing rewilding to restoration – Rejecting the search for novelty

    Hayward, M. W., Scanlon, R. J., Callen, A., Howell, L. G., Klop-Toker, K. L., Di Blanco, Y., Balkenhol, N., Bugir, C. K., Campbell, L., Caravaggi, A., Chalmers, A. C., Clulow, J., Clulow, S., Cross, P., Gould, J. A., Griffin, A. S., Heurich, M., Howe, B. K., Jachowski, D. S., Jhala, Y. V., Krishnamurthy, R., Kowalczyk, R., Lenga, D. J., Linnell, J. D. C., Marnewick, K. A., Moehrenschlager, A., Montgomery, R. A., Osipova, L., Peneaux, C., Rodger, J. C., Sales, L. P., Seeto, R. G. Y., Shuttleworth, C. M., Somers, M. J., Tamessar, C. T., Upton, R. M. O. & Weise, F. J., 1 May 2019, In: Biological Conservation. 233, p. 255-259 5 p.

    Research output: Contribution to journalArticlepeer-review

  32. Published

    Deconstructing compassionate conservation

    Hayward, M. W., Callen, A., Allen, B. L., Ballard, G., Broekhuis, F., Bugir, C., Clarke, R. H., Clulow, J., Clulow, S., Daltry, J. C., Davies-Mostert, H. T., Fleming, P. J. S., Griffin, A. S., Howell, L. G., Kerley, G. I. H., Klop-Toker, K., Legge, S., Major, T., Meyer, N., Montgomery, R. A., Moseby, K., Parker, D. M., Périquet, S., Read, J., Scanlon, R., Seeto, R., Shuttleworth, C., Somers, M. J., Tamessar, C. T., Tuft, K., Upton, R., Valenzuela-Molina, M., Wayne, A., Witt, R. R. & Wuster, W., Aug 2019, In: Conservation Biology. 33, 4, p. 760-768

    Research output: Contribution to journalArticlepeer-review

  33. Published

    Repeated squirrelpox infection in Welsh red squirrels

    Holmes, J. P., Everest, D. & Shuttleworth, C., 3 Feb 2023, In: Veterinary Record. 192, 3, p. 128-129

    Research output: Contribution to journalLetter

  34. Published

    First report of dermatophilosis in wild European red squirrels (Sciurus vulgaris)

    Holmes, P., Everest, D. J., Spiro, S., Wessels, M. & Shuttleworth, C., 30 Sept 2019, In: Veterinary Record Case Reports. 7, 3, p. 3

    Research output: Contribution to journalArticlepeer-review

  35. Published

    By trying to be everything, rewilding risks meaning nothing

    Jachowski, J. D., Bombaci, S., Caravaggi, A., Chalmers, A., Clulow, S., Griffin, A., Hayward, M., Heurich, M., Linnell, J., Marnewick, K., Moehrenschlager, A., Montgomery, A., Shuttleworth, C., Somers, M. & Weise, F., Nov 2019, In: Science. 364, p. 6438

    Research output: Contribution to journalLetter

  36. Published

    Mathematical models of grey squirrel invasion: a case study on Anglesey

    Jones, H. E., White, A., Lurz, P. W. W., Boots, M. & Shuttleworth, C., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (eds.). European Squirrel Initiative, p. 235-255

    Research output: Chapter in Book/Report/Conference proceedingChapterpeer-review

  37. Published

    Mathematical models for invasive species management: grey squirrel control on Anglesey

    Jones, H., White, A., Lurz, P. W. W. & Shuttleworth, C., 10 Sept 2017, In: Ecological Modelling. 180, p. 274-284

    Research output: Contribution to journalArticlepeer-review

  38. Published

    Are novel ecosystems the only novelty of rewilding?

    Klop-Toker, K., Clulow, S., Shuttleworth, C. & Hayward, M., 1 Nov 2020, In: Restoration Ecology. 28, p. 1318-1320

    Research output: Contribution to journalArticlepeer-review

  39. Published

    Ranging behaviour, density and social structure in grey squirrels

    Lawton, C., Shuttleworth, C. & Kenward, R. E., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (eds.). European Squirrel Initiative, p. 133-152

    Research output: Chapter in Book/Report/Conference proceedingChapterpeer-review

  40. Published

    Reintroductions and translocations of red squirrels within Europe

    Lawton, C., Waters, C. & Shuttleworth, C., 2015, Red squirrels: Ecology, Conservation & Management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Hayward, M. W. (eds.). European Squirrel Initiative, p. 193-210

    Research output: Chapter in Book/Report/Conference proceedingChapterpeer-review

  41. Published

    Introduced Canadian Eastern grey squirrels: squirrelpox virus surveillance and why nothing matters.

    McInnes, C., Shuttleworth, C., Larsen, K. W., Everest, D. J., Bruemmer, C., Carroll, B., Romeo, C., Sainsbury, A. W., Crawshaw, G., Dubois, S., Gillis, L., Gilray, J. & Percival, A., 3 Aug 2020, In: Hystrix-Italian Journal of Mammalogy. 31, p. 95-98

    Research output: Contribution to journalArticlepeer-review

  42. Published

    Genetic management of the red squirrel, Sciurus vulgaris: a practical approach to regional conservation

    Ogden, R., Shuttleworth, C., McEwing, R. & Cesarini, S., 26 Oct 2005, In: Conservation Genetics. 6, p. 511-525

    Research output: Contribution to journalArticlepeer-review

  43. Published

    Ethical dilemmas when using citizen science for early detection of invasive tree pests and diseases

    Pocock, M. J. O., Marzano, M., Bullas-Appleton, E., Dyke, A., de Groot, M., Shuttleworth, C. & White, R., 1 Nov 2020, In: Management of Biological Invasions. 11, p. 720-732

    Research output: Contribution to journalArticlepeer-review

  44. Published

    The large-scale removal of mammalian invasive alien species in Northern Europe

    Robertson, P. A., Adriaens, T., Lambin, X., Mill, A., Roy, S., Shuttleworth, C. & Sutton-Croft, M., 1 Feb 2017, In: Pest Management Science. 73, p. 273-279

    Research output: Contribution to journalArticlepeer-review

  45. Published

    Invasive alien species colonisation prevention: Your guide to early detection and rapid response

    Robinson, N. & Shuttleworth, C., 2020, Newark: Royal Society of Wildlife Trusts. 204 p.

    Research output: Book/ReportBookpeer-review

  46. Published

    A practical guide to early surveillance and monitoring

    Robinson, N. J., Everest, D. J., Harry, P. & Shuttleworth, C., 2021, Invasive Alien Species Colonisation Prevention: Your guide to early detection and rapid response . Robinson, N. J. & Shuttleworth, C. M. (eds.). Royal Society of Wildlife Trusts, p. 163-202

    Research output: Chapter in Book/Report/Conference proceedingChapter

  47. Published

    Disease, invasions and conservation: no evidence of squirrelpox virus in grey squirrels introduced to Italy

    Romeo, C., McInnes, C., Dale, T., Shuttleworth, C., Bertolino, S., Wauters, L. A. & Ferrari, N., 1 Feb 2019, In: Animal Conservation. 22, 1, p. 14-23

    Research output: Contribution to journalArticlepeer-review

  48. Published

    British red squirrels remain the only known wild rodent host for leprosy bacilli

    Schilling, A., Avanzi, C., Ulrich, R. G., Busso, P., Pisanu, B., Ferrari, N., Romeo, C., Mazzamuto, M. V., McLuckie, J., Shuttleworth, C., Del-Pozo, J., Lurz, P. W. W., Escalante-Fuentes, W. G., Ocampo-Candiani, J., Vera-Cabrera, L., Stevenson, K., Chapuis, J-L., Meredith, A. L. & Cole, S. T., 1 Feb 2019, In: Frontiers in Veterinary Science. 6, p. 8

    Research output: Contribution to journalArticlepeer-review

  49. Published

    Leprosy in red squirrels in the UK

    Schilling, A-K., Del-Pozo, J., Lurz, P. W. W., Stevenson, K., Avanzi, C., Shuttleworth, C., Cole, S. T. & Meredith, A. L., 30 Mar 2019, In: Veterinary Record. 184, p. 416

    Research output: Contribution to journalArticle

  50. Published

    Landscape scale impacts of culling upon a European grey squirrel population: can trapping reduce population size and decrease the threat of squirrel pox virus infection for the native red squirrel?

    Schuchert, P., Shuttleworth, C., McInnes, C. J., Everest, D. J. & Rushton, S. P., 1 Nov 2014, In: Biological Invasions. 16, p. 2381-2391

    Research output: Contribution to journalArticlepeer-review

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