Research Portal

Dr Craig Shuttleworth

Honorary Visiting Research Fellow

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ORCID: 0000-0003-0076-6789

Contact info

Mobile 07966150847

Email c.shuttleworth@bangor.ac.uk

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  1. Article › Research › Not peer-reviewed
  2. Published

    Adenovirus detections in juvenile squirrels

    Everest, D. J., Dastjerdi, A., Cowan, D., Gomm, M., Shuttleworth, C., McInnes, C. J., Deane, D., Coulter, L., Mill, A., Rushton, S. P., Jackson, N. L. & Litherland, P., 1 Oct 2015, In: Veterinary Record. 177, p. 373-374

    Research output: Contribution to journal › Article

  3. Published

    Adenovirus infection in red squirrels

    Shuttleworth, C., Everest, D. J., Stidworthy, M., Halliwell, E. C., Clews-Roberts, R., Paine, V., Bamber, C. & Mason, J., 31 Dec 2021, In: The Veterinary record. 189, p. 444-445

    Research output: Contribution to journal › Article

  4. Published

    Adenovirus infection in red squirrels in areas free from grey squirrels

    Everest, D. J., Butler, H., Simpson, V. R., Blackett, T. & Shuttleworth, C., 31 Aug 2013, In: The Veterinary record. 173, 8, p. 199-200

    Research output: Contribution to journal › Article

  5. Published

    Confirmation of squirrelpox in Welsh red squirrel

    Everest, D. J., Floyd, T., Donnnacie, B., Irvine, R. M., Holmes, J. P. & Shuttleworth, C., 30 Nov 2017, In: Veterinary Record. 181, p. 514-515

    Research output: Contribution to journal › Article

  6. Published

    Detecting viral infection in red squirrels

    Shuttleworth, C., Everest, D. J., Halliwell, E. C., Hulme, B., Wilberforce, L. & Clews-Roberts, R., 30 Apr 2019, In: Veterinary Record. 184, p. 507

    Research output: Contribution to journal › Article

  7. Published

    Leprosy in red squirrels in the UK

    Schilling, A-K., Del-Pozo, J., Lurz, P. W. W., Stevenson, K., Avanzi, C., Shuttleworth, C., Cole, S. T. & Meredith, A. L., 30 Mar 2019, In: Veterinary Record. 184, p. 416

    Research output: Contribution to journal › Article

  8. Published

    Non-invasive genetic studies and virus epidemiology

    Everest, D. J., Dastjerdi, A., Davies, H., Shuttleworth, C., Ogden, R., Butler, H. & Sapsford, B., 30 Mar 2022, In: Veterinary Record. 190, p. 204-205

    Research output: Contribution to journal › Article

  9. Published

    Pine marten scat holds few clues for squirrel disease

    Shuttleworth, C., Everest, D. J., Dastjerdi, A., Birks, JDS. & Hayward, M., 7 Dec 2019, In: Veterinary Record. 185, p. 698

    Research output: Contribution to journal › Article

  10. Published

    Red squirrel conservation and disease - Adenovirus in Wales

    Shuttleworth, C., 2014, In: Natur Cymru. 52, p. 41-44

    Research output: Contribution to journal › Article

  11. Published

    Red squirrels on Anglesey

    Shuttleworth, C. & Bailey, M., 2003, In: Natur Cymru. 6, p. 35-37

    Research output: Contribution to journal › Article

  12. Published

    Releasing grey squirrels into the wild

    Shuttleworth, C., Bertolino, S., Gill, R., Gurnell, J., Hayward, M., Kenward, R. E., Lawton, C., Lurz, P. W. W., McInnes, C. J., Mill, A., Trotter, S. & Wauters, L. A., 31 Mar 2019, In: Veterinary Record. 184, p. 389-390

    Research output: Contribution to journal › Article

  13. Published

    Turning the grey tide: progress in red squirrel recovery

    Shuttleworth, C., 1 Jun 2010, In: Ecos. 31, p. 27-35

    Research output: Contribution to journal › Article

  14. Published

    Virus presence within Welsh red squirrels

    Everest, D. J., Holmes, P. & Shuttleworth, C., 30 May 2017, In: The Veterinary record. 180, p. 520-521

    Research output: Contribution to journal › Article

  15. Article › Research › Peer-reviewed
  16. Published

    A preliminary comparison of magnum 116 body-grip traps with a live capture trap design during the control of grey squirrels (Sciurus carolinensis)

    Shuttleworth, C., Mill, A. & Van der Waal, Z., Dec 2016, In: International Journal of Pest Management. 63, 4, p. 316-321

    Research output: Contribution to journal › Article › peer-review

  17. Published

    A study of nest box use by a low density red squirrel (Sciurus vulgaris) population within a commercial conifer plantation

    Shuttleworth, C., 2004, In: Quarterly Journal of Forestry. 98, p. 113-119

    Research output: Contribution to journal › Article › peer-review

  18. Published

    A systematic assessment of the impact of adenovirus infection on a captive re-introduction project for red squirrels (Sciurus vulgaris)

    Everest, D. J., Shuttleworth, C., Grierson, S. S., Duff, J. P., Jackson, N., Litherland, P. & Kenward, R. E., 31 Aug 2012, In: Veterinary Record. 171, p. 176-184

    Research output: Contribution to journal › Article › peer-review

  19. Published

    A temporal refuge from predation can change the outcome of prey species competition

    Slade, A., White, A., Lurz, P. W. W., Shuttleworth, C. & Lambin, X., 1 Sept 2022, In: Oikos. Oikos e08565

    Research output: Contribution to journal › Article › peer-review

  20. Published

    A tough nut to crack: red squirrel conservation in Wales

    Shuttleworth, C., 1 Oct 2003, In: Biologist. 50, p. 231-235

    Research output: Contribution to journal › Article › peer-review

  21. Published

    Adenovirus infection: An emerging factor in red squirrel conservation programmes

    Everest, D. J., Shuttleworth, C., Stidworthy, M. F., Grierson, S. S., Duff, P. J. & Kenward, R. E., 1 Oct 2014, In: Mammal Review. 44, 3-4, p. 225-233

    Research output: Contribution to journal › Article › peer-review

  22. Published

    Adenovirus-associated deaths in red squirrels on Anglesey

    Everest, D. J., Shuttleworth, C. & Stidworthy, M. F., 31 Oct 2008, In: The Veterinary record. 163, 14, p. 430

    Research output: Contribution to journal › Article › peer-review

  23. Published

    Against a grey tide: red squirrel conservation on Anglesey

    Shuttleworth, C., 2005, In: Quarterly Journal of Forestry. 99, p. 213-214

    Research output: Contribution to journal › Article › peer-review

  24. Published

    An Opportunistic Assessment of the Impact of Squirrelpox Disease Outbreaks upon a Red Squirrel Population Sympatric with Grey Squirrels in Wales

    Shuttleworth, C., Everest, D., Holmes, P., Bell, S. & Cripps, R., 1 Jan 2022, In: Animals. 12, 1

    Research output: Contribution to journal › Article › peer-review

  25. Published

    Are nest boxes a useful tool in regional red squirrel conservation programs?

    Shuttleworth, C. & Schuchert, P., 15 Dec 2014, In: Hystrix-Italian Journal of Mammalogy. 25, p. 91-94

    Research output: Contribution to journal › Article › peer-review

  26. Published

    Are novel ecosystems the only novelty of rewilding?

    Klop-Toker, K., Clulow, S., Shuttleworth, C. & Hayward, M., 1 Nov 2020, In: Restoration Ecology. 28, p. 1318-1320

    Research output: Contribution to journal › Article › peer-review

  27. Published

    Assessing a potential non-invasive method for viral diagnostic purposes in European squirrels

    Everest, D. J., Tolhurst-Cherriman, D. A. R., Davies, H., Dastjerdi, A., Ashton, A., Blackett, T., Meredith, A. L., Milne, E., Mill, A. & Shuttleworth, C., 4 Jun 2019, In: Hystrix-Italian Journal of Mammalogy. 30, p. 44-50

    Research output: Contribution to journal › Article › peer-review

  28. Published

    Assessing causes and significance of red squirrel (Sciurus vulgaris) mortality during regional population restoration: An applied conservation perspective

    Shuttleworth, C., Signorile, A. L., Everest, D. J., Lurz, P. W. W. & Duff, J. P., 16 Nov 2015, In: Hystrix-Italian Journal of Mammalogy. 26, 2, p. 69-75

    Research output: Contribution to journal › Article › peer-review

  29. Published

    British red squirrels remain the only known wild rodent host for leprosy bacilli

    Schilling, A., Avanzi, C., Ulrich, R. G., Busso, P., Pisanu, B., Ferrari, N., Romeo, C., Mazzamuto, M. V., McLuckie, J., Shuttleworth, C., Del-Pozo, J., Lurz, P. W. W., Escalante-Fuentes, W. G., Ocampo-Candiani, J., Vera-Cabrera, L., Stevenson, K., Chapuis, J-L., Meredith, A. L. & Cole, S. T., 1 Feb 2019, In: Frontiers in Veterinary Science. 6, p. 8

    Research output: Contribution to journal › Article › peer-review

  30. Published

    Capturing red squirrels (Sciurus vulgaris) on camera: A cost-effective approach for monitoring relative abundance and habitat preference

    Shannon, G., Valle, S. & Shuttleworth, C., 3 Oct 2023, In: Ecology and Evolution. 13, 10, e10536.

    Research output: Contribution to journal › Article › peer-review

  31. Published

    Challenges and solutions in early detection, rapid response and communication about potential invasive alien species in forests

    de Groot, M., Bullas-Appleton, E., Csóka, G., Csiszár, A., Faccoli, A., Gervasini, A., Kirichenko, N., Shuttleworth, C., Kus Veenvliet, J., O'Hanlon, R., Korda, M., Marinsek, A., Robinson, N., Sweeney, J., Tricarico, E., Verbrugge, L., Williams, D. & Zidar, S., 14 Aug 2020, In: Management of Biological Invasions. 11, p. 637-660

    Research output: Contribution to journal › Article › peer-review

  32. Published

    Deconstructing compassionate conservation

    Hayward, M. W., Callen, A., Allen, B. L., Ballard, G., Broekhuis, F., Bugir, C., Clarke, R. H., Clulow, J., Clulow, S., Daltry, J. C., Davies-Mostert, H. T., Fleming, P. J. S., Griffin, A. S., Howell, L. G., Kerley, G. I. H., Klop-Toker, K., Legge, S., Major, T., Meyer, N., Montgomery, R. A., Moseby, K., Parker, D. M., Périquet, S., Read, J., Scanlon, R., Seeto, R., Shuttleworth, C., Somers, M. J., Tamessar, C. T., Tuft, K., Upton, R., Valenzuela-Molina, M., Wayne, A., Witt, R. R. & Wuster, W., Aug 2019, In: Conservation Biology. 33, 4, p. 760-768

    Research output: Contribution to journal › Article › peer-review

  33. Published

    Disease monitoring and surveillance: case studies in the applied conservation of fragmented red squirrel (Sciurus vulgaris) populations in England and Wales

    Everest, D. J., Floyd, T., Holmes, P., Man, C., Dunnett, C., Locke, R., Savage, L., Sutcliffe, S., Sapsford, B., Shuttleworth, C. & Duff, P., 1 Dec 2021, In: Mammalian Biology. 101, p. 1079-1088

    Research output: Contribution to journal › Article › peer-review

  34. Published

    Disease, invasions and conservation: no evidence of squirrelpox virus in grey squirrels introduced to Italy

    Romeo, C., McInnes, C., Dale, T., Shuttleworth, C., Bertolino, S., Wauters, L. A. & Ferrari, N., 1 Feb 2019, In: Animal Conservation. 22, 1, p. 14-23

    Research output: Contribution to journal › Article › peer-review

  35. Published

    Do differing levels of boldness influence the success of translocation? A pilot study on red squirrels (Sciurus vulgaris)

    Bamber, J. A., Shuttleworth, C. & Hayward, M., 26 Sept 2020, In: Animals. 10, 1748.

    Research output: Contribution to journal › Article › peer-review

  36. Published

    Envisioning the future with 'compassionate conservation': An ominous projection for native wildlife and biodiversity

    Callen, A., Hayward, M. W., Klop-Toker, K., Allen, B. L., Ballard, G., Beranek, C. T., Broekhuis, F., Bugir, C. K., Clarke, R. H., Clulow, J., Clulow, S., Daltry, J. C., Davies-Mostert, H. T., Di Blanco, Y. E., Dixon, V., Fleming, P. J. S., Howell, L. G., Kerley, G. I. H., Legge, S. M., Lenga, D. J., Major, T., Montgomery, R. A., Moseby, K., Meyer, N., Parker, D. M., Periquet, S., Read, J., Scanlon, R. J., Shuttleworth, C., Tamessar, C. T., Taylor, W. A., Tuft, K., Upton, R. M. O., Valenzuela, M., Witt, R. R. & Wuster, W., Jan 2020, In: Biological Conservation. 241, 108365.

    Research output: Contribution to journal › Article › peer-review

  37. Published

    Ethical dilemmas when using citizen science for early detection of invasive tree pests and diseases

    Pocock, M. J. O., Marzano, M., Bullas-Appleton, E., Dyke, A., de Groot, M., Shuttleworth, C. & White, R., 1 Nov 2020, In: Management of Biological Invasions. 11, p. 720-732

    Research output: Contribution to journal › Article › peer-review

  38. Published

    Evolving grey squirrel management techniques in Europe

    Shuttleworth, C., Robinson, N. J., Halliwell, E. C., Clews-Roberts, R., Peek, H., Podgornik, G., Stinson, M., Rice, S., Finlay, C., McKinney, C., Everest, D. J. & Larsen, K. W., 21 Oct 2020, In: Management of Biological Invasions. 11, 4, p. 747-761

    Research output: Contribution to journal › Article › peer-review

  39. Published

    First report of dermatophilosis in wild European red squirrels (Sciurus vulgaris)

    Holmes, P., Everest, D. J., Spiro, S., Wessels, M. & Shuttleworth, C., 30 Sept 2019, In: Veterinary Record Case Reports. 7, 3, p. 3

    Research output: Contribution to journal › Article › peer-review

  40. Published

    Genetic management of the red squirrel, Sciurus vulgaris: a practical approach to regional conservation

    Ogden, R., Shuttleworth, C., McEwing, R. & Cesarini, S., 26 Oct 2005, In: Conservation Genetics. 6, p. 511-525

    Research output: Contribution to journal › Article › peer-review

  41. Published

    Indirect effects of pine marten recovery result in benefits to native prey through suppression of an invasive species and a shared pathogen

    Slade, A., White, A., Lurz, P. W. W., Shuttleworth, C., Tosh, D. & Twining, JP., 1 Feb 2023, In: Ecological Modelling. 476, 110216.

    Research output: Contribution to journal › Article › peer-review

  42. Published

    Integrating red squirrel (Sciurus vulgaris) habitat requirements with the management of pathogenic tree disease in commercial forests in the UK

    Shuttleworth, C., Lurz, P. W. W., Geddes, N. & Brown, J., 1 Sept 2012, In: Forest Ecology and Management. 279, p. 167-175

    Research output: Contribution to journal › Article › peer-review

  43. Published

    Inter-specific viral infections: Can the management of captive red squirrel collections help inform scientific research?

    Shuttleworth, C., Everest, D. J., McInnes, C. J., Greenwood, A., Jackson, N., Rushton, S. P. & Kenward, R. E., 16 Jun 2014, In: Hystrix-Italian Journal of Mammalogy. 25, 1, p. 18-24

    Research output: Contribution to journal › Article › peer-review

  44. Published

    Interactions between the red squirrel (Sciurus vulgaris), great tit (Parus Major) and jackdaw (Corvus monedula) whilst using nest boxes.

    Shuttleworth, C., 1 Oct 2001, In: Journal of Zoology. 255, p. 269-272

    Research output: Contribution to journal › Article › peer-review

  45. Published

    Introduced Canadian Eastern grey squirrels: squirrelpox virus surveillance and why nothing matters.

    McInnes, C., Shuttleworth, C., Larsen, K. W., Everest, D. J., Bruemmer, C., Carroll, B., Romeo, C., Sainsbury, A. W., Crawshaw, G., Dubois, S., Gillis, L., Gilray, J. & Percival, A., 3 Aug 2020, In: Hystrix-Italian Journal of Mammalogy. 31, p. 95-98

    Research output: Contribution to journal › Article › peer-review

  46. E-pub ahead of print

    Invasive genetic rescue: dispersal following repeated culling reinforces the genetic diversity of an invasive mammal

    Synnott, R., Shuttleworth, C., Everest, D., Stevenson-Holt, C., O'Reilly, C., McDevitt, A. & O.Meara, D., 30 Jun 2023, (E-pub ahead of print) In: Biological Invasions.

    Research output: Contribution to journal › Article › peer-review

  47. Published

    Landscape scale impacts of culling upon a European grey squirrel population: can trapping reduce population size and decrease the threat of squirrel pox virus infection for the native red squirrel?

    Schuchert, P., Shuttleworth, C., McInnes, C. J., Everest, D. J. & Rushton, S. P., 1 Nov 2014, In: Biological Invasions. 16, p. 2381-2391

    Research output: Contribution to journal › Article › peer-review

  48. Published

    Mathematical models for invasive species management: grey squirrel control on Anglesey

    Jones, H., White, A., Lurz, P. W. W. & Shuttleworth, C., 10 Sept 2017, In: Ecological Modelling. 180, p. 274-284

    Research output: Contribution to journal › Article › peer-review

  49. Published

    Opportunistic viral surveillance confirms the ongoing disease threat grey squirrels pose to sympatric red squirrel populations in the UK

    Everest, DJ., Green, C., Dastjerdi, A., Davies, H., Cripps, R., McKinney, C., Podgornik, G., Stinson, M., O'Hare, S., Bonnie, S., Mill, A., Van der wall, Z., Robinson, N., Trotter, S. & Shuttleworth, C., 20 May 2023, In: Veterinary Record. 192, 10, e2834.

    Research output: Contribution to journal › Article › peer-review

  50. Published

    PCR detection of adenovirus in grey squirrels on Anglesey

    Everest, D. J., Grierson, S. S., Stidworthy, M. & Shuttleworth, C., 31 Oct 2009, In: Veterinary Record. 165, p. 482

    Research output: Contribution to journal › Article › peer-review

  51. Published

    Recalibrating Risk: Implications of squirrelpox virus for successful red squirrel translocations within mainland UK

    Shuttleworth, C., Brady, D., Cross, P., Gardner, L., Greenwood, A., Jackson, N., McKinney, C., Robinson, N., Trotter, S., Valle, S., Wood, K. & Hayward, M., Feb 2021, In: Conservation Science and Practice. 3, 2, e321.

    Research output: Contribution to journal › Article › peer-review

  52. Published

    Red squirrel bark-stripping of hornbeam in North Wales

    Shuttleworth, C. & Gill, R., 2019, In: Quarterly Journal of Forestry. 113, 4, p. 275-277

    Research output: Contribution to journal › Article › peer-review

  53. Published

    Reducing negative impacts on non-target species during grey squirrel (Sciurus carolinensis) control activities

    Shuttleworth, C., Mill, A. C., Van der Wall, Z., Bertolino, S. & Everest, D. J., 1 Jan 2018, In: Quarterly Journal of Forestry. 112, 1, p. 21-29

    Research output: Contribution to journal › Article › peer-review

  54. Published

    Reinstating trophic cascades as an applied conservation tool to protect forest ecosystems from invasive grey squirrels (Sciurus carolinensis)

    Bamber, J. A., Shuttleworth, C. M., Hayward, M. W. & Everest, D. J., 31 Dec 2020, In: Food Webs. 25, e00164.

    Research output: Contribution to journal › Article › peer-review

  55. Published

    Reintroducing rewilding to restoration – Rejecting the search for novelty

    Hayward, M. W., Scanlon, R. J., Callen, A., Howell, L. G., Klop-Toker, K. L., Di Blanco, Y., Balkenhol, N., Bugir, C. K., Campbell, L., Caravaggi, A., Chalmers, A. C., Clulow, J., Clulow, S., Cross, P., Gould, J. A., Griffin, A. S., Heurich, M., Howe, B. K., Jachowski, D. S., Jhala, Y. V., Krishnamurthy, R., Kowalczyk, R., Lenga, D. J., Linnell, J. D. C., Marnewick, K. A., Moehrenschlager, A., Montgomery, R. A., Osipova, L., Peneaux, C., Rodger, J. C., Sales, L. P., Seeto, R. G. Y., Shuttleworth, C. M., Somers, M. J., Tamessar, C. T., Upton, R. M. O. & Weise, F. J., 1 May 2019, In: Biological Conservation. 233, p. 255-259 5 p.

    Research output: Contribution to journal › Article › peer-review

  56. Retrospective detection by negative contrast electron microscopy of faecal viral particles in wild red squirrels (Sciurus vulgaris) with suspected enteropathy in Great Britain

    Everest, D. J., Stidworthy, M. F., Milne, E. M., Meredith, A. L., Chantery, J., Shuttleworth, C., Blackett, T., Butler, H., Wilkinson, M. & Sainsbury, A. W., 31 Dec 2010, In: Veterinary Record. 167, p. 1007-1010

    Research output: Contribution to journal › Article › peer-review

  57. Published

    Significant differences in the caecal bacterial microbiota of red and grey squirrels in Britain

    Hall, L., Nichols, C., Leng, J., Martelli, F., Shuttleworth, C. & La Ragione, R., 14 Feb 2024, In: Journal of Medical Microbiology. 73, 2

    Research output: Contribution to journal › Article › peer-review

  58. Published

    Sympatric cases of squirrelpox in red and grey squirrels

    Shuttleworth, C., Everest, D., Wood, C. & Holmes, J. P., 7 May 2021, In: The Veterinary record. 188, 9, p. 358

    Research output: Contribution to journal › Article › peer-review

  59. Published

    The foraging behaviour and diet of red squirrels (Sciurus vulgaris) receiving supplemental feeding

    Shuttleworth, C., 1 Sept 2000, In: Wildlife Biology. 6, p. 149-156

    Research output: Contribution to journal › Article › peer-review

  60. Published

    The implications of significant adenovirus infection in UK captive red squirrel (Sciurus vulgaris) collections: How histological screening can aid applied conservation management

    Everest, D. J., Shuttleworth, C., Grierson, S. S., Dastjerdi, A., M.F., S., Duff, J. P., Higgins, R. J., Mill, A. C. & Chantrey, J., 1 Jan 2018, In: Mammalian Biology. 88, p. 123-129

    Research output: Contribution to journal › Article › peer-review

  61. Published

    The large-scale removal of mammalian invasive alien species in Northern Europe

    Robertson, P. A., Adriaens, T., Lambin, X., Mill, A., Roy, S., Shuttleworth, C. & Sutton-Croft, M., 1 Feb 2017, In: Pest Management Science. 73, p. 273-279

    Research output: Contribution to journal › Article › peer-review

  62. Published

    The search for novelty continues for rewilding

    Hayward, M., Jachowski, J. D., Bugir, C. J., Clulow, J., Krishnamurthy, R., Griffin, A. S., Chalmers, A. C., Linnell, J. D. C., Montgomery, R. A., Somers, M. J., Kowalczyk, R., Heurich, M., Caravaggi, A., Marnewick, K. A., Di Blanco, Y., Shuttleworth, C., Callen, C., Weise, F., Scanlon, R., Moehrenschlager, A., Howell, L. & Upton, R. M. O., 1 Aug 2019, In: Biological Conservation. 236, p. 584-585

    Research output: Contribution to journal › Article › peer-review

  63. Published

    The use of nest boxes by the red squirrel (Sciurus vulgaris) in conifer plantations

    Shuttleworth, C., 2001, In: Quarterly Journal of Forestry. 95, p. 225-228

    Research output: Contribution to journal › Article › peer-review

  64. Published

    The use of nest boxes by the red squirrel Sciurus vulgaris in a coniferous habitat

    Shuttleworth, C., 1 Mar 1999, In: Mammal Review. 29, p. 61-66

    Research output: Contribution to journal › Article › peer-review

  65. Published

    Traffic related mortality in a red squirrel (Sciurus vulgaris) population receiving supplemental feeding

    Shuttleworth, C., 1 Jun 2001, In: Urban Ecosystems. 2, p. 109-118

    Research output: Contribution to journal › Article › peer-review

  66. Letter › Research › Not peer-reviewed
  67. Published

    Adenovirus remains widespread in squirrels on Anglesey

    Shuttleworth, C., Everest, D. & McQueen, A., 16 Dec 2023, In: Veterinary Record. 193, 12, p. 490-491 2 p.

    Research output: Contribution to journal › Letter

  68. Published

    By trying to be everything, rewilding risks meaning nothing

    Jachowski, J. D., Bombaci, S., Caravaggi, A., Chalmers, A., Clulow, S., Griffin, A., Hayward, M., Heurich, M., Linnell, J., Marnewick, K., Moehrenschlager, A., Montgomery, A., Shuttleworth, C., Somers, M. & Weise, F., Nov 2019, In: Science. 364, p. 6438

    Research output: Contribution to journal › Letter

  69. Published

    Repeated squirrelpox infection in Welsh red squirrels

    Holmes, J. P., Everest, D. & Shuttleworth, C., 3 Feb 2023, In: Veterinary Record. 192, 3, p. 128-129

    Research output: Contribution to journal › Letter

  70. Published

    Rewilding needs to clarify the role of management of invasive species

    Hayward, M., Jachowski, D. S., Shuttleworth, C., Linnell, J. D. C., Allen, B. L., Griffen, A. S., Montgomery, R. A., Caravaggi, A., Weise, F. J., Moehrenschlager, A., Somers, M. J., Clulow, S., Heurich, M. & Marnewick, K., 8 May 2019, In: Science. 364, p. 6438

    Research output: Contribution to journal › Letter

  71. Letter › Research › Peer-reviewed
  72. Published

    Response to comments on "Compassionate Conservation deserves a morally serious rather than dismissive response - reply to Callen et al., 2020"

    Callen, A., Hayward, M. W., Klop-Toker, K., Allen, B. L., Ballard, G., Beranek, C. T., Broekhuis, F., Bugir, C. K., Clarke, R. H., Clulow, J., Clulow, S., Daltry, J. C., Davies-Mostert, H. T., Di Blanco, Y. E., Dixon, V., Fleming, P. J. S., Howell, L. G., Kerley, G. I. H., Legge, S. M., Lenga, D. J., Major, T., Montgomery, R. A., Moseby, K., Meyer, N., Parker, D. M., Periquet, S., Read, J., Scanlon, R. J., Shuttleworth, C., Tamessar, C. T., Taylor, W. A., Tuft, K., Upton, R. M. O., Valenzuela, M., Witt, R. R. & Wuster, W., 30 Apr 2020, In: Biological Conservation. 244

    Research output: Contribution to journal › Letter › peer-review

  73. Chapter › Research › Not peer-reviewed
  74. Published

    A practical guide to early surveillance and monitoring

    Robinson, N. J., Everest, D. J., Harry, P. & Shuttleworth, C., 2021, Invasive Alien Species Colonisation Prevention: Your guide to early detection and rapid response . Robinson, N. J. & Shuttleworth, C. M. (eds.). Royal Society of Wildlife Trusts, p. 163-202

    Research output: Chapter in Book/Report/Conference proceeding › Chapter

  75. Published

    Against a grey tide: What future for the red squirrel?

    Shuttleworth, C., 2005, Journal of Practical Ecology & Conservation (Special Series No 4): Living with the Aliens? - Ecology, History, Culture and Management if exotic plants and animals: issues for nature conservation . Rotherham, L. D. (ed.). Vol. 4. p. 87-88

    Research output: Chapter in Book/Report/Conference proceeding › Chapter

  76. Published

    Developing red squirrel re-introduction techniques for use during regional grey squirrel eradication programmes in Europe

    Shuttleworth, C., Jackson, N. & Kenward, R. E., 2015, 2015 Global Reintroduction Perspectives. Soorae, P. (ed.). IUCN/SSC Re-introduction Specialist Group, p. 67-74

    Research output: Chapter in Book/Report/Conference proceeding › Chapter

  77. Published

    Island Haven: Anglesey red squirrel conservation

    Shuttleworth, C., 2015, Shared Experience of Red Squirrel Conservation Practice . Shuttleworth, C. M., Lurz, P. W. W. & Halliwell, E. C. (eds.). European Squirrel Initiative, p. 151-158

    Research output: Chapter in Book/Report/Conference proceeding › Chapter

  78. Published

    Magical Mammals: Red squirrels in North West Wales

    Shuttleworth, C., 2021, Saving the red squirrel: landscape recovery. Shuttleworth, C. M., Lurz, P. W. W. & Robinson, N. J. (eds.). Red Squirrel Survival Trust, p. 269-278

    Research output: Chapter in Book/Report/Conference proceeding › Chapter

  79. Published

    Monitoring escaped captive bred red squirrels in the UK

    Shuttleworth, C., 2021, Saving the red squirrel: landscape recovery. Shuttleworth, C. M., Lurz, P. W. W. & Robinson, N. J. (eds.). Red Squirrel Survival Trust, p. 71-78

    Research output: Chapter in Book/Report/Conference proceeding › Chapter

  80. Published

    Painting the Town Red: Gwynedd red squirrel conservation

    Shuttleworth, C., 2015, Shared Experience of Red Squirrel Conservation Practice . Shuttleworth, C. M., Lurz, P. W. W. & Halliwell, E. C. (eds.). European Squirrel Initiative, p. 159-166

    Research output: Chapter in Book/Report/Conference proceeding › Chapter

  81. Published

    Preventing grey squirrel establishment on Anglesey

    Shuttleworth, C., 2021, Invasive Alien Species Colonisation Prevention: Your guide to early detection and rapid response . Robinson, N. J. & Shuttleworth, C. M. (eds.). Royal Society of Wildlife Trusts, p. 55-69

    Research output: Chapter in Book/Report/Conference proceeding › Chapter

  82. Published

    Re-introduction of the red squirrel into Newborough forest on the island of Anglesey, UK

    Shuttleworth, C., Kenward, R. E. & Jackson, N., 2008, Global Re-Introduction Perspectives: Re-Introduction Case-Studies from Around the Globe. Soorae, P. S. (ed.). IUCN/SSC Re-introduction Specialist Group, p. 163-166

    Research output: Chapter in Book/Report/Conference proceeding › Chapter

  83. Published

    The management of coastal sand dune woodland for red squirrels (Sciurus vulgaris L.)

    Shuttleworth, C. & Gurnell, J., 2001, Coastal Dune Management: Shared Experience of European Conservation Practice. Houston, J., Edmondson, A. S. E. & Rooney, P. J. (eds.). Liverpool University Press, p. 117-127

    Research output: Chapter in Book/Report/Conference proceeding › Chapter

  84. Chapter › Research › Peer-reviewed
  85. Published

    British red squirrel strongholds: challenges for conservation

    Gurnell, J., Blackett, T., Butler, H., Lurz, P. W. W., Magris, L. & Shuttleworth, C., 2015, Red squirrels: Ecology, Conservation & Management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Hayward, M. W. (eds.). European Squirrel Initiative, p. 211-232

    Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review

  86. Published

    Developing integrated and applied red squirrel conservation programmes: What lessons can Europe learn from a regional grey squirrel eradication programme in North Wales?

    Shuttleworth, C., Schuchert, P., Everest, D. J., McInnes, C. J., Rushton, S. P., Jackson, N. & Kenward, R. E., 2015, Red squirrels: Ecology, Conservation & Management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Hayward, M. W. (eds.). European Squirrel Initiative, p. 233-250

    Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review

  87. Published

    Ecosystem impacts of an alien invader in Europe, the grey squirrel Sciurus carolinensis

    Gurnell, J., Lurz, P. W. W. & Shuttleworth, C., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (eds.). European Squirrel Initiative, p. 307-329

    Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review

  88. Published

    Experiences of grey squirrel management in an upland conifer forest

    Halliwell, E. C., Shuttleworth, C., Cartmel, S., Lloyd, I. & Jenkins, R., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (eds.). European Squirrel Initiative, p. 453-474

    Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review

  89. Published

    Genetic evidence of the effectiveness of grey squirrel control operations: lessons from the Isle of Anglesey

    Signorile, L. & Shuttleworth, C., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (eds.). European Squirrel Initiative, p. 439-452

    Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review

  90. Published

    Grey squirrel control as a tactic in the conservation of the red squirrel: a positive outcome

    Shuttleworth, C., 2001, Advances in Vertebrate Pest Management . Feare, C. J. & Cowan, D. P. (eds.). Filander Verlag, Vol. 3. p. 151-168

    Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review

  91. Published

    Grey squirrel management in Europe - the future

    Shuttleworth, C., Lurz, P. W. W. & Gurnell, J., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (eds.). European Squirrel Initiative

    Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review

  92. Published

    Grey squirrel nesting ecology and the use of nest sites in European population management

    Shuttleworth, C., Selonen, V. & Koprowski, J. L., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (eds.). European Squirrel Initiative, p. 349-369

    Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review

  93. Published

    Identifying incursion pathways, early detection responses and management actions to prevent grey squirrel range expansion: an island case study in Wales

    Shuttleworth, C., Halliwell, E. C. & Robertson, P. A., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (eds.). European Squirrel Initiative, p. 475-494

    Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review

  94. Published

    Mathematical models of grey squirrel invasion: a case study on Anglesey

    Jones, H. E., White, A., Lurz, P. W. W., Boots, M. & Shuttleworth, C., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (eds.). European Squirrel Initiative, p. 235-255

    Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review

  95. Published

    Ranging behaviour, density and social structure in grey squirrels

    Lawton, C., Shuttleworth, C. & Kenward, R. E., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (eds.). European Squirrel Initiative, p. 133-152

    Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review

  96. Published

    Red squirrels (Sciurus vulgaris) on the Sefton Coast: The conservation value of nest boxes

    Shuttleworth, C., 1995, 2nd NPI Red Alert UK Forum for Red Squirrel Conservation. Hughes, D. G. & Tew, T. (eds.). The Federation of Zoological Gardens of Great Britain and Ireland, p. 25-32

    Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review

  97. Published

    Reintroductions and translocations of red squirrels within Europe

    Lawton, C., Waters, C. & Shuttleworth, C., 2015, Red squirrels: Ecology, Conservation & Management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Hayward, M. W. (eds.). European Squirrel Initiative, p. 193-210

    Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review

  98. Published

    Striving for success: an evaluation of local action to conserve red squirrels (Sciurus vulgaris) in Wales

    Halliwell, E. C., Shuttleworth, C., Wilberforce, L., Denman, H., Lloyd, I. & Cartmel, S., 2015, Red squirrels: Ecology, Conservation & Management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Hayward, M. W. (eds.). European Squirrel Initiative, p. 175-192

    Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review

  99. Published

    The effect of supplemental feeding on the diet, population density and reproduction of red squirrels

    Shuttleworth, C., 1997, The Conservation of Red Squirrels (Sciurus vulgaris L.). Gurnell, J. & Lurz, P. W. W. (eds.). Peoples Trust for Endangered Species, p. 13-24

    Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review

  100. Published

    The impact of adenoviruses on red squirrel (Sciurus vulgaris) conservation – with notes on other enteric viruses of potential significance

    Everest, D. J., Shuttleworth, C., Grierson, S. S., Dastjerdi, A., Stidworthy, M. F., Duff, J. P. & Sainsbury, A. W., 2015, Red squirrels: Ecology, Conservation & Management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Hayward, M. W. (eds.). European Squirrel Initiative, p. 129-146

    Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review

  101. Published

    The management of grey squirrel populations in Europe: evolving best practice

    Bertolino, S., Lurz, P. W. W., Shuttleworth, C., Martinoli, A. & Wauters, L. A., 2016, The grey squirrel: ecology and management in Europe . Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (eds.). European Squirrel Initiative, p. 495-516

    Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review

  102. Published

    The potential role of ectoparasites in the epidemiology of squirrelpox virus: a possible novel means of intervention to reduce the impact of the disease?

    Cowan, D. P., Mill, A. C., Everest, D. J., McInnes, C. J., Rushton, S. P., Shirley, M. D. F., Start, C. & Shuttleworth, C., 2016, The grey squirrel: ecology and management in Europe . Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (eds.). European Squirrel Initiative, p. 255-289

    Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review

  103. Published

    The role of spruce in red squirrel (Sciurus vulgaris) conservation in the UK

    Shuttleworth, C. & Lurz, P. W. W., 2014, Forest Ecosystems: Biodiversity, Management and Conservation. Roberts, N. C. (ed.). Nova Science Publishers, p. 63-78

    Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review

  104. Article › Research
  105. Published

    Gut bacteria may explain why grey squirrels outcompete reds – new research

    Shuttleworth, C., 20 Feb 2024, The Conversation.

    Research output: Contribution to specialist publication › Article

  106. Other contribution › Research › Not peer-reviewed
  107. Published

    Grey squirrels are bad for the British countryside – full stop

    Shuttleworth, C., 2017, The Conversation.

    Research output: Other contribution

  108. Published

    Rare woodland wildlife at risk because of 50-year-old tree felling rules

    Shuttleworth, C., 2018, The Conversation.

    Research output: Other contribution

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