Dr Craig Shuttleworth
Honorary Visiting Research Fellow

Affiliations
Links
- https://twitter.com/Welshsquirrels
Personal account - https://twitter.com/RedSquirrelsinW
Welsh red squirrel conservation - https://www.facebook.com/groups/143610382316300
Welsh red squirrel conservation - https://pinemarten.net/
The Gwynedd pine marten project - https://www.linkedin.com/in/craig-shuttleworth-6062a1253/
Personal page
Contact info
Mobile 07966150847
Email c.shuttleworth@bangor.ac.uk
- 2021
- Published
Saving the red squirrel: landscape scale recovery
Shuttleworth, C., Lurz, P. W. W. & Robinson, N., 2021, London: Red Squirrel Survival Trust. 306 p.Research output: Book/Report › Book › peer-review
- Published
Recalibrating Risk: Implications of squirrelpox virus for successful red squirrel translocations within mainland UK
Shuttleworth, C., Brady, D., Cross, P., Gardner, L., Greenwood, A., Jackson, N., McKinney, C., Robinson, N., Trotter, S., Valle, S., Wood, K. & Hayward, M., Feb 2021, In: Conservation Science and Practice. 3, 2, e321.Research output: Contribution to journal › Article › peer-review
- Published
Sympatric cases of squirrelpox in red and grey squirrels
Shuttleworth, C., Everest, D., Wood, C. & Holmes, J. P., 7 May 2021, In: The Veterinary record. 188, 9, p. 358Research output: Contribution to journal › Article › peer-review
- Published
Disease monitoring and surveillance: case studies in the applied conservation of fragmented red squirrel (Sciurus vulgaris) populations in England and Wales
Everest, D. J., Floyd, T., Holmes, P., Man, C., Dunnett, C., Locke, R., Savage, L., Sutcliffe, S., Sapsford, B., Shuttleworth, C. & Duff, P., 1 Dec 2021, In: Mammalian Biology. 101, p. 1079-1088Research output: Contribution to journal › Article › peer-review
- Published
Adenovirus infection in red squirrels
Shuttleworth, C., Everest, D. J., Stidworthy, M., Halliwell, E. C., Clews-Roberts, R., Paine, V., Bamber, C. & Mason, J., 31 Dec 2021, In: The Veterinary record. 189, p. 444-445Research output: Contribution to journal › Article
- 2022
- Published
An Opportunistic Assessment of the Impact of Squirrelpox Disease Outbreaks upon a Red Squirrel Population Sympatric with Grey Squirrels in Wales
Shuttleworth, C., Everest, D., Holmes, P., Bell, S. & Cripps, R., 1 Jan 2022, In: Animals. 12, 1Research output: Contribution to journal › Article › peer-review
- Published
Non-invasive genetic studies and virus epidemiology
Everest, D. J., Dastjerdi, A., Davies, H., Shuttleworth, C., Ogden, R., Butler, H. & Sapsford, B., 30 Mar 2022, In: Veterinary Record. 190, p. 204-205Research output: Contribution to journal › Article
- Published
A temporal refuge from predation can change the outcome of prey species competition
Slade, A., White, A., Lurz, P. W. W., Shuttleworth, C. & Lambin, X., 1 Sept 2022, In: Oikos. Oikos e08565Research output: Contribution to journal › Article › peer-review
- Published
Squirrelpox outbreak detected in north Wales – without a vaccine, the disease will keep decimating red squirrels
Shuttleworth, C., 21 Dec 2022, The Conversation.Research output: Other contribution
- 2023
- Published
Restoring our lost wildlife: a review of the ecological, social and economic factors in the potential reintroduction of six mammal species to Cornwall
Cooper, P., Brazier, R., Puttock, A., Graham, H., Auster, R., Campbell-Palmer, R., Shuttleworth, C. & Pizzi, R., 2023, 258 p.Research output: Book/Report › Commissioned report
- Published
Indirect effects of pine marten recovery result in benefits to native prey through suppression of an invasive species and a shared pathogen
Slade, A., White, A., Lurz, P. W. W., Shuttleworth, C., Tosh, D. & Twining, J., 1 Feb 2023, In: Ecological Modelling. 476, 110216.Research output: Contribution to journal › Article › peer-review
- Published
Repeated squirrelpox infection in Welsh red squirrels
Holmes, J. P., Everest, D. & Shuttleworth, C., 3 Feb 2023, In: Veterinary Record. 192, 3, p. 128-129Research output: Contribution to journal › Letter
- Published
Opportunistic viral surveillance confirms the ongoing disease threat grey squirrels pose to sympatric red squirrel populations in the UK
Everest, D., Green, C., Dastjerdi, A., Davies, H., Cripps, R., McKinney, C., Podgornik, G., Stinson, M., O'Hare, S., Bonnie, S., Mill, A., Van der wall, Z., Robinson, N., Trotter, S. & Shuttleworth, C., 20 May 2023, In: Veterinary Record. 192, 10, e2834.Research output: Contribution to journal › Article › peer-review
- Published
Invasive genetic rescue: dispersal following repeated culling reinforces the genetic diversity of an invasive mammal
Synnott, R., Shuttleworth, C., Everest, D., Stevenson-Holt, C., O'Reilly, C., McDevitt, A. & O.Meara, D., Oct 2023, In: Biological Invasions. 25, p. 3323-3339Research output: Contribution to journal › Article › peer-review
- Published
Capturing red squirrels (Sciurus vulgaris) on camera: A cost-effective approach for monitoring relative abundance and habitat preference
Shannon, G., Valle, S. & Shuttleworth, C., 3 Oct 2023, In: Ecology and Evolution. 13, 10, e10536.Research output: Contribution to journal › Article › peer-review
- Published
Adenovirus remains widespread in squirrels on Anglesey
Shuttleworth, C., Everest, D. & McQueen, A., 16 Dec 2023, In: Veterinary Record. 193, 12, p. 490-491 2 p.Research output: Contribution to journal › Letter
- 2024
- Published
Significant differences in the caecal bacterial microbiota of red and grey squirrels in Britain
Hall, L., Nichols, C., Leng, J., Martelli, F., Shuttleworth, C. & La Ragione, R., 14 Feb 2024, In: Journal of Medical Microbiology. 73, 2Research output: Contribution to journal › Article › peer-review
- Published
Gut bacteria may explain why grey squirrels outcompete reds – new research
Shuttleworth, C., 20 Feb 2024, The Conversation.Research output: Contribution to specialist publication › Article
- Published
Developing a method for assessing the relative abundance of red squirrels in low-density populations
Shannon, G., Barton, O., Valle, S. & Shuttleworth, C., 1 Nov 2024, 34 p.Research output: Book/Report › Commissioned report
- 2025
- Published
Mitochondrial DNA Analysis of Red Squirrels (Sciurus vulgaris) in North Wales
O’Meara, D., Shuttleworth, C., Clews-Roberts, R., Everest, D. & Synnott, R., 1 Mar 2025, 14 p. (Evidence Report Series; no. 791)Research output: Book/Report › Commissioned report