Dr Craig Shuttleworth

Honorary Visiting Research Fellow

Contact info

Mobile 07966150847

Email c.shuttleworth@bangor.ac.uk

  1. Published
  2. Published

    Grey squirrel management in Europe - the future

    Shuttleworth, C., Lurz, P. W. W. & Gurnell, J., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (eds.). European Squirrel Initiative

    Research output: Chapter in Book/Report/Conference proceedingChapterpeer-review

  3. Published

    Monitoring escaped captive bred red squirrels in the UK

    Shuttleworth, C., 2021, Saving the red squirrel: landscape recovery. Shuttleworth, C. M., Lurz, P. W. W. & Robinson, N. J. (eds.). Red Squirrel Survival Trust, p. 71-78

    Research output: Chapter in Book/Report/Conference proceedingChapter

  4. Published

    Red squirrels (Sciurus vulgaris) on the Sefton Coast: The conservation value of nest boxes

    Shuttleworth, C., 1995, 2nd NPI Red Alert UK Forum for Red Squirrel Conservation. Hughes, D. G. & Tew, T. (eds.). The Federation of Zoological Gardens of Great Britain and Ireland, p. 25-32

    Research output: Chapter in Book/Report/Conference proceedingChapterpeer-review

  5. Published

    Red squirrel bark-stripping of hornbeam in North Wales

    Shuttleworth, C. & Gill, R., 2019, In: Quarterly Journal of Forestry. 113, 4, p. 275-277

    Research output: Contribution to journalArticlepeer-review

  6. Published

    Red squirrels: ecology, conservation & management in Europe

    Shuttleworth, C., Hayward, M. & Lurz, P. W. W., 2015, European Squirrel Initiative. 330 p.

    Research output: Book/ReportBookpeer-review

  7. Published

    Pine marten scat holds few clues for squirrel disease

    Shuttleworth, C., Everest, D. J., Dastjerdi, A., Birks, J. & Hayward, M., 7 Dec 2019, In: Veterinary Record. 185, p. 698

    Research output: Contribution to journalArticle

  8. Published

    Recalibrating Risk: Implications of squirrelpox virus for successful red squirrel translocations within mainland UK

    Shuttleworth, C., Brady, D., Cross, P., Gardner, L., Greenwood, A., Jackson, N., McKinney, C., Robinson, N., Trotter, S., Valle, S., Wood, K. & Hayward, M., Feb 2021, In: Conservation Science and Practice. 3, 2, e321.

    Research output: Contribution to journalArticlepeer-review

  9. Published

    Integrating red squirrel (Sciurus vulgaris) habitat requirements with the management of pathogenic tree disease in commercial forests in the UK

    Shuttleworth, C., Lurz, P. W. W., Geddes, N. & Brown, J., 1 Sept 2012, In: Forest Ecology and Management. 279, p. 167-175

    Research output: Contribution to journalArticlepeer-review

  10. Published

    The use of nest boxes by the red squirrel Sciurus vulgaris in a coniferous habitat

    Shuttleworth, C., 1 Mar 1999, In: Mammal Review. 29, p. 61-66

    Research output: Contribution to journalArticlepeer-review

  11. Published

    Grey squirrel nesting ecology and the use of nest sites in European population management

    Shuttleworth, C., Selonen, V. & Koprowski, J. L., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (eds.). European Squirrel Initiative, p. 349-369

    Research output: Chapter in Book/Report/Conference proceedingChapterpeer-review

  12. Published

    Why the pine marten is not every red squirrel’s best friend

    Shuttleworth, C. & Hayward, M., 2019, The Conversation.

    Research output: Other contribution

  13. Published

    The use of nest boxes by the red squirrel (Sciurus vulgaris) in conifer plantations

    Shuttleworth, C., 2001, In: Quarterly Journal of Forestry. 95, p. 225-228

    Research output: Contribution to journalArticlepeer-review

  14. Published

    An Opportunistic Assessment of the Impact of Squirrelpox Disease Outbreaks upon a Red Squirrel Population Sympatric with Grey Squirrels in Wales

    Shuttleworth, C., Everest, D., Holmes, P., Bell, S. & Cripps, R., 1 Jan 2022, In: Animals. 12, 1

    Research output: Contribution to journalArticlepeer-review

  15. Published

    Developing red squirrel re-introduction techniques for use during regional grey squirrel eradication programmes in Europe

    Shuttleworth, C., Jackson, N. & Kenward, R. E., 2015, 2015 Global Reintroduction Perspectives. Soorae, P. (ed.). IUCN/SSC Re-introduction Specialist Group, p. 67-74

    Research output: Chapter in Book/Report/Conference proceedingChapter

  16. Published

    Rare woodland wildlife at risk because of 50-year-old tree felling rules

    Shuttleworth, C., 2018, The Conversation.

    Research output: Other contribution

  17. Published
  18. Published

    Capturing red squirrels (Sciurus vulgaris) on camera: A cost-effective approach for monitoring relative abundance and habitat preference

    Shannon, G., Valle, S. & Shuttleworth, C., 3 Oct 2023, In: Ecology and Evolution. 13, 10, e10536.

    Research output: Contribution to journalArticlepeer-review

  19. Published
  20. Published

    Landscape scale impacts of culling upon a European grey squirrel population: can trapping reduce population size and decrease the threat of squirrel pox virus infection for the native red squirrel?

    Schuchert, P., Shuttleworth, C., McInnes, C. J., Everest, D. J. & Rushton, S. P., 1 Nov 2014, In: Biological Invasions. 16, p. 2381-2391

    Research output: Contribution to journalArticlepeer-review

  21. Published

    British red squirrels remain the only known wild rodent host for leprosy bacilli

    Schilling, A., Avanzi, C., Ulrich, R. G., Busso, P., Pisanu, B., Ferrari, N., Romeo, C., Mazzamuto, M. V., McLuckie, J., Shuttleworth, C., Del-Pozo, J., Lurz, P. W. W., Escalante-Fuentes, W. G., Ocampo-Candiani, J., Vera-Cabrera, L., Stevenson, K., Chapuis, J.-L., Meredith, A. L. & Cole, S. T., 1 Feb 2019, In: Frontiers in Veterinary Science. 6, p. 8

    Research output: Contribution to journalArticlepeer-review

  22. Published

    Leprosy in red squirrels in the UK

    Schilling, A.-K., Del-Pozo, J., Lurz, P. W. W., Stevenson, K., Avanzi, C., Shuttleworth, C., Cole, S. T. & Meredith, A. L., 30 Mar 2019, In: Veterinary Record. 184, p. 416

    Research output: Contribution to journalArticle

  23. Published

    Disease, invasions and conservation: no evidence of squirrelpox virus in grey squirrels introduced to Italy

    Romeo, C., McInnes, C., Dale, T., Shuttleworth, C., Bertolino, S., Wauters, L. A. & Ferrari, N., 1 Feb 2019, In: Animal Conservation. 22, 1, p. 14-23

    Research output: Contribution to journalArticlepeer-review

  24. Published

    Invasive alien species colonisation prevention: Your guide to early detection and rapid response

    Robinson, N. & Shuttleworth, C., 2020, Newark: Royal Society of Wildlife Trusts. 204 p.

    Research output: Book/ReportBookpeer-review

  25. Published

    A practical guide to early surveillance and monitoring

    Robinson, N. J., Everest, D. J., Harry, P. & Shuttleworth, C., 2021, Invasive Alien Species Colonisation Prevention: Your guide to early detection and rapid response . Robinson, N. J. & Shuttleworth, C. M. (eds.). Royal Society of Wildlife Trusts, p. 163-202

    Research output: Chapter in Book/Report/Conference proceedingChapter

  26. Published

    The large-scale removal of mammalian invasive alien species in Northern Europe

    Robertson, P. A., Adriaens, T., Lambin, X., Mill, A., Roy, S., Shuttleworth, C. & Sutton-Croft, M., 1 Feb 2017, In: Pest Management Science. 73, p. 273-279

    Research output: Contribution to journalArticlepeer-review

  27. Published

    Ethical dilemmas when using citizen science for early detection of invasive tree pests and diseases

    Pocock, M. J. O., Marzano, M., Bullas-Appleton, E., Dyke, A., de Groot, M., Shuttleworth, C. & White, R., 1 Nov 2020, In: Management of Biological Invasions. 11, p. 720-732

    Research output: Contribution to journalArticlepeer-review

  28. Published

    Mitochondrial DNA Analysis of Red Squirrels (Sciurus vulgaris) in North Wales

    O’Meara, D., Shuttleworth, C., Clews-Roberts, R., Everest, D. & Synnott, R., 1 Mar 2025, 14 p. (Evidence Report Series; no. 791)

    Research output: Book/ReportCommissioned report

  29. Published

    Genetic management of the red squirrel, Sciurus vulgaris: a practical approach to regional conservation

    Ogden, R., Shuttleworth, C., McEwing, R. & Cesarini, S., 26 Oct 2005, In: Conservation Genetics. 6, p. 511-525

    Research output: Contribution to journalArticlepeer-review

  30. Published

    Introduced Canadian Eastern grey squirrels: squirrelpox virus surveillance and why nothing matters.

    McInnes, C., Shuttleworth, C., Larsen, K. W., Everest, D. J., Bruemmer, C., Carroll, B., Romeo, C., Sainsbury, A. W., Crawshaw, G., Dubois, S., Gillis, L., Gilray, J. & Percival, A., 3 Aug 2020, In: Hystrix-Italian Journal of Mammalogy. 31, p. 95-98

    Research output: Contribution to journalArticlepeer-review

  31. Published

    Ranging behaviour, density and social structure in grey squirrels

    Lawton, C., Shuttleworth, C. & Kenward, R. E., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (eds.). European Squirrel Initiative, p. 133-152

    Research output: Chapter in Book/Report/Conference proceedingChapterpeer-review

  32. Published

    Reintroductions and translocations of red squirrels within Europe

    Lawton, C., Waters, C. & Shuttleworth, C., 2015, Red squirrels: Ecology, Conservation & Management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Hayward, M. W. (eds.). European Squirrel Initiative, p. 193-210

    Research output: Chapter in Book/Report/Conference proceedingChapterpeer-review

  33. Published

    Are novel ecosystems the only novelty of rewilding?

    Klop-Toker, K., Clulow, S., Shuttleworth, C. & Hayward, M., 1 Nov 2020, In: Restoration Ecology. 28, p. 1318-1320

    Research output: Contribution to journalArticlepeer-review

  34. Published

    Mathematical models of grey squirrel invasion: a case study on Anglesey

    Jones, H. E., White, A., Lurz, P. W. W., Boots, M. & Shuttleworth, C., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (eds.). European Squirrel Initiative, p. 235-255

    Research output: Chapter in Book/Report/Conference proceedingChapterpeer-review

  35. Published

    Mathematical models for invasive species management: grey squirrel control on Anglesey

    Jones, H., White, A., Lurz, P. W. W. & Shuttleworth, C., 10 Sept 2017, In: Ecological Modelling. 180, p. 274-284

    Research output: Contribution to journalArticlepeer-review

  36. Published

    By trying to be everything, rewilding risks meaning nothing

    Jachowski, J. D., Bombaci, S., Caravaggi, A., Chalmers, A., Clulow, S., Griffin, A., Hayward, M., Heurich, M., Linnell, J., Marnewick, K., Moehrenschlager, A., Montgomery, A., Shuttleworth, C., Somers, M. & Weise, F., Nov 2019, In: Science. 364, p. 6438

    Research output: Contribution to journalLetter

  37. Published

    Repeated squirrelpox infection in Welsh red squirrels

    Holmes, J. P., Everest, D. & Shuttleworth, C., 3 Feb 2023, In: Veterinary Record. 192, 3, p. 128-129

    Research output: Contribution to journalLetter

  38. Published

    First report of dermatophilosis in wild European red squirrels (Sciurus vulgaris)

    Holmes, P., Everest, D. J., Spiro, S., Wessels, M. & Shuttleworth, C., 30 Sept 2019, In: Veterinary Record Case Reports. 7, 3, p. 3

    Research output: Contribution to journalArticlepeer-review

  39. Published

    Rewilding needs to clarify the role of management of invasive species

    Hayward, M., Jachowski, D. S., Shuttleworth, C., Linnell, J. D. C., Allen, B. L., Griffen, A. S., Montgomery, R. A., Caravaggi, A., Weise, F. J., Moehrenschlager, A., Somers, M. J., Clulow, S., Heurich, M. & Marnewick, K., 8 May 2019, In: Science. 364, p. 6438

    Research output: Contribution to journalLetter

  40. Published

    The search for novelty continues for rewilding

    Hayward, M., Jachowski, J. D., Bugir, C. J., Clulow, J., Krishnamurthy, R., Griffin, A. S., Chalmers, A. C., Linnell, J. D. C., Montgomery, R. A., Somers, M. J., Kowalczyk, R., Heurich, M., Caravaggi, A., Marnewick, K. A., Di Blanco, Y., Shuttleworth, C., Callen, C., Weise, F., Scanlon, R., Moehrenschlager, A., Howell, L. & Upton, R. M. O., 1 Aug 2019, In: Biological Conservation. 236, p. 584-585

    Research output: Contribution to journalArticlepeer-review

  41. Published

    Reintroducing rewilding to restoration – Rejecting the search for novelty

    Hayward, M. W., Scanlon, R. J., Callen, A., Howell, L. G., Klop-Toker, K. L., Di Blanco, Y., Balkenhol, N., Bugir, C. K., Campbell, L., Caravaggi, A., Chalmers, A. C., Clulow, J., Clulow, S., Cross, P., Gould, J. A., Griffin, A. S., Heurich, M., Howe, B. K., Jachowski, D. S., Jhala, Y. V., Krishnamurthy, R., Kowalczyk, R., Lenga, D. J., Linnell, J. D. C., Marnewick, K. A., Moehrenschlager, A., Montgomery, R. A., Osipova, L., Peneaux, C., Rodger, J. C., Sales, L. P., Seeto, R. G. Y., Shuttleworth, C. M., Somers, M. J., Tamessar, C. T., Upton, R. M. O. & Weise, F. J., 1 May 2019, In: Biological Conservation. 233, p. 255-259 5 p.

    Research output: Contribution to journalArticlepeer-review

  42. Published

    Deconstructing compassionate conservation

    Hayward, M. W., Callen, A., Allen, B. L., Ballard, G., Broekhuis, F., Bugir, C., Clarke, R. H., Clulow, J., Clulow, S., Daltry, J. C., Davies-Mostert, H. T., Fleming, P. J. S., Griffin, A. S., Howell, L. G., Kerley, G. I. H., Klop-Toker, K., Legge, S., Major, T., Meyer, N., Montgomery, R. A., Moseby, K., Parker, D. M., Périquet, S., Read, J., Scanlon, R., Seeto, R., Shuttleworth, C., Somers, M. J., Tamessar, C. T., Tuft, K., Upton, R., Valenzuela-Molina, M., Wayne, A., Witt, R. R. & Wuster, W., Aug 2019, In: Conservation Biology. 33, 4, p. 760-768

    Research output: Contribution to journalArticlepeer-review

  43. Published

    Striving for success: an evaluation of local action to conserve red squirrels (Sciurus vulgaris) in Wales

    Halliwell, E. C., Shuttleworth, C., Wilberforce, L., Denman, H., Lloyd, I. & Cartmel, S., 2015, Red squirrels: Ecology, Conservation & Management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Hayward, M. W. (eds.). European Squirrel Initiative, p. 175-192

    Research output: Chapter in Book/Report/Conference proceedingChapterpeer-review

  44. Published

    Experiences of grey squirrel management in an upland conifer forest

    Halliwell, E. C., Shuttleworth, C., Cartmel, S., Lloyd, I. & Jenkins, R., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (eds.). European Squirrel Initiative, p. 453-474

    Research output: Chapter in Book/Report/Conference proceedingChapterpeer-review

  45. Published

    Significant differences in the caecal bacterial microbiota of red and grey squirrels in Britain

    Hall, L., Nichols, C., Leng, J., Martelli, F., Shuttleworth, C. & La Ragione, R., 14 Feb 2024, In: Journal of Medical Microbiology. 73, 2

    Research output: Contribution to journalArticlepeer-review

  46. Published

    British red squirrel strongholds: challenges for conservation

    Gurnell, J., Blackett, T., Butler, H., Lurz, P. W. W., Magris, L. & Shuttleworth, C., 2015, Red squirrels: Ecology, Conservation & Management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Hayward, M. W. (eds.). European Squirrel Initiative, p. 211-232

    Research output: Chapter in Book/Report/Conference proceedingChapterpeer-review

  47. Published

    Ecosystem impacts of an alien invader in Europe, the grey squirrel Sciurus carolinensis

    Gurnell, J., Lurz, P. W. W. & Shuttleworth, C., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (eds.). European Squirrel Initiative, p. 307-329

    Research output: Chapter in Book/Report/Conference proceedingChapterpeer-review

  48. Published

    Controlling grey squirrels in forests and woodlands in the UK

    Gill, R., Ferryman, M., Shuttleworth, C., Lurz, P. W. W., Mill, A., Robertson, P. A. & Dutton, C., 2019, Forest Research. 16 p.

    Research output: Book/ReportBookpeer-review

  49. Published

    Non-invasive genetic studies and virus epidemiology

    Everest, D. J., Dastjerdi, A., Davies, H., Shuttleworth, C., Ogden, R., Butler, H. & Sapsford, B., 30 Mar 2022, In: Veterinary Record. 190, p. 204-205

    Research output: Contribution to journalArticle

  50. Published

    Adenovirus infection in red squirrels in areas free from grey squirrels

    Everest, D. J., Butler, H., Simpson, V. R., Blackett, T. & Shuttleworth, C., 31 Aug 2013, In: The Veterinary record. 173, 8, p. 199-200

    Research output: Contribution to journalArticle