Research Portal

Dr Craig Shuttleworth

Honorary Visiting Research Fellow

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ORCID: 0000-0003-0076-6789

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Mobile 07966150847

Email c.shuttleworth@bangor.ac.uk

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  1. Published

    Grey squirrel nesting ecology and the use of nest sites in European population management

    Shuttleworth, C., Selonen, V. & Koprowski, J. L., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (eds.). European Squirrel Initiative, p. 349-369

    Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review

  2. Published

    Why the pine marten is not every red squirrel’s best friend

    Shuttleworth, C. & Hayward, M., 2019, The Conversation.

    Research output: Other contribution

  3. Published

    Turning the grey tide: progress in red squirrel recovery

    Shuttleworth, C., 1 Jun 2010, In: Ecos. 31, p. 27-35

    Research output: Contribution to journal › Article

  4. Published

    Red squirrel conservation and disease - Adenovirus in Wales

    Shuttleworth, C., 2014, In: Natur Cymru. 52, p. 41-44

    Research output: Contribution to journal › Article

  5. Published

    Shared experience of red squirrel conservation practice

    Shuttleworth, C., Lurz, P. W. W. & Halliwell, E. C., 2015, European Squirrel Initiative. 220 p.

    Research output: Book/Report › Book

  6. Published

    Red squirrels (Sciurus vulgaris) on the Sefton Coast: The conservation value of nest boxes

    Shuttleworth, C., 1995, 2nd NPI Red Alert UK Forum for Red Squirrel Conservation. Hughes, D. G. & Tew, T. (eds.). The Federation of Zoological Gardens of Great Britain and Ireland, p. 25-32

    Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review

  7. Published

    Are nest boxes a useful tool in regional red squirrel conservation programs?

    Shuttleworth, C. & Schuchert , P., 15 Dec 2014, In: Hystrix-Italian Journal of Mammalogy. 25, p. 91-94

    Research output: Contribution to journal › Article › peer-review

  8. Published

    Re-introduction of the red squirrel into Newborough forest on the island of Anglesey, UK

    Shuttleworth, C., Kenward, R. E. & Jackson, N., 2008, Global Re-Introduction Perspectives: Re-Introduction Case-Studies from Around the Globe. Soorae, P. S. (ed.). IUCN/SSC Re-introduction Specialist Group, p. 163-166

    Research output: Chapter in Book/Report/Conference proceeding › Chapter

  9. Published

    Painting the Town Red: Gwynedd red squirrel conservation

    Shuttleworth, C., 2015, Shared Experience of Red Squirrel Conservation Practice . Shuttleworth, C. M., Lurz, P. W. W. & Halliwell, E. C. (eds.). European Squirrel Initiative, p. 159-166

    Research output: Chapter in Book/Report/Conference proceeding › Chapter

  10. Published

    Reducing negative impacts on non-target species during grey squirrel (Sciurus carolinensis) control activities

    Shuttleworth, C., Mill, A. C., Van der Wall, Z., Bertolino, S. & Everest, D. J., 1 Jan 2018, In: Quarterly Journal of Forestry. 112, 1, p. 21-29

    Research output: Contribution to journal › Article › peer-review

  11. Published

    Red squirrels in my garden: guidance and tips to help conserve local populations

    Shuttleworth, C. & Halliwell, E. C., 2018, European Squirrel Initiative. 185 p.

    Research output: Book/Report › Book › peer-review

  12. Published

    Against a grey tide: What future for the red squirrel?

    Shuttleworth, C., 2005, Journal of Practical Ecology & Conservation (Special Series No 4): Living with the Aliens? - Ecology, History, Culture and Management if exotic plants and animals: issues for nature conservation . Rotherham, L. D. (ed.). Vol. 4. p. 87-88

    Research output: Chapter in Book/Report/Conference proceeding › Chapter

  13. Published

    A tough nut to crack: red squirrel conservation in Wales

    Shuttleworth, C., 1 Oct 2003, In: Biologist. 50, p. 231-235

    Research output: Contribution to journal › Article › peer-review

  14. Published

    Traffic related mortality in a red squirrel (Sciurus vulgaris) population receiving supplemental feeding

    Shuttleworth, C., 1 Jun 2001, In: Urban Ecosystems. 2, p. 109-118

    Research output: Contribution to journal › Article › peer-review

  15. Published

    An Opportunistic Assessment of the Impact of Squirrelpox Disease Outbreaks upon a Red Squirrel Population Sympatric with Grey Squirrels in Wales

    Shuttleworth, C., Everest, D., Holmes, P., Bell, S. & Cripps, R., 1 Jan 2022, In: Animals. 12, 1

    Research output: Contribution to journal › Article › peer-review

  16. Published

    Island Haven: Anglesey red squirrel conservation

    Shuttleworth, C., 2015, Shared Experience of Red Squirrel Conservation Practice . Shuttleworth, C. M., Lurz, P. W. W. & Halliwell, E. C. (eds.). European Squirrel Initiative, p. 151-158

    Research output: Chapter in Book/Report/Conference proceeding › Chapter

  17. Published

    Rare woodland wildlife at risk because of 50-year-old tree felling rules

    Shuttleworth, C., 2018, The Conversation.

    Research output: Other contribution

  18. Published

    Capturing red squirrels (Sciurus vulgaris) on camera: A cost-effective approach for monitoring relative abundance and habitat preference

    Shannon, G., Valle, S. & Shuttleworth, C., 3 Oct 2023, In: Ecology and Evolution. 13, 10, e10536.

    Research output: Contribution to journal › Article › peer-review

  19. Published
  20. Published

    Landscape scale impacts of culling upon a European grey squirrel population: can trapping reduce population size and decrease the threat of squirrel pox virus infection for the native red squirrel?

    Schuchert, P., Shuttleworth, C., McInnes, C. J., Everest, D. J. & Rushton, S. P., 1 Nov 2014, In: Biological Invasions. 16, p. 2381-2391

    Research output: Contribution to journal › Article › peer-review

  21. Published

    Leprosy in red squirrels in the UK

    Schilling, A.-K., Del-Pozo, J., Lurz, P. W. W., Stevenson, K., Avanzi, C., Shuttleworth, C., Cole, S. T. & Meredith, A. L., 30 Mar 2019, In: Veterinary Record. 184, p. 416

    Research output: Contribution to journal › Article

  22. Published

    British red squirrels remain the only known wild rodent host for leprosy bacilli

    Schilling, A., Avanzi, C., Ulrich, R. G., Busso, P., Pisanu, B., Ferrari, N., Romeo, C., Mazzamuto, M. V., McLuckie, J., Shuttleworth, C., Del-Pozo, J., Lurz, P. W. W., Escalante-Fuentes, W. G., Ocampo-Candiani, J., Vera-Cabrera, L., Stevenson, K., Chapuis, J.-L., Meredith, A. L. & Cole, S. T., 1 Feb 2019, In: Frontiers in Veterinary Science. 6, p. 8

    Research output: Contribution to journal › Article › peer-review

  23. Published

    Disease, invasions and conservation: no evidence of squirrelpox virus in grey squirrels introduced to Italy

    Romeo, C., McInnes, C., Dale, T., Shuttleworth, C., Bertolino, S., Wauters, L. A. & Ferrari, N., 1 Feb 2019, In: Animal Conservation. 22, 1, p. 14-23

    Research output: Contribution to journal › Article › peer-review

  24. Published

    Invasive alien species colonisation prevention: Your guide to early detection and rapid response

    Robinson, N. & Shuttleworth, C., 2020, Newark: Royal Society of Wildlife Trusts. 204 p.

    Research output: Book/Report › Book › peer-review

  25. Published

    A practical guide to early surveillance and monitoring

    Robinson, N. J., Everest, D. J., Harry, P. & Shuttleworth, C., 2021, Invasive Alien Species Colonisation Prevention: Your guide to early detection and rapid response . Robinson, N. J. & Shuttleworth, C. M. (eds.). Royal Society of Wildlife Trusts, p. 163-202

    Research output: Chapter in Book/Report/Conference proceeding › Chapter

  26. Published

    The large-scale removal of mammalian invasive alien species in Northern Europe

    Robertson, P. A., Adriaens, T., Lambin, X., Mill, A., Roy, S., Shuttleworth, C. & Sutton-Croft, M., 1 Feb 2017, In: Pest Management Science. 73, p. 273-279

    Research output: Contribution to journal › Article › peer-review

  27. Published

    Ethical dilemmas when using citizen science for early detection of invasive tree pests and diseases

    Pocock, M. J. O., Marzano, M., Bullas-Appleton, E., Dyke, A., de Groot, M., Shuttleworth, C. & White, R., 1 Nov 2020, In: Management of Biological Invasions. 11, p. 720-732

    Research output: Contribution to journal › Article › peer-review

  28. Published

    Mitochondrial DNA Analysis of Red Squirrels (Sciurus vulgaris) in North Wales

    O’Meara, D., Shuttleworth, C., Clews-Roberts, R., Everest, D. & Synnott, R., 1 Mar 2025, 14 p. (Evidence Report Series; no. 791)

    Research output: Book/Report › Commissioned report

  29. Published

    Genetic management of the red squirrel, Sciurus vulgaris: a practical approach to regional conservation

    Ogden, R., Shuttleworth, C., McEwing, R. & Cesarini, S., 26 Oct 2005, In: Conservation Genetics. 6, p. 511-525

    Research output: Contribution to journal › Article › peer-review

  30. Published

    Introduced Canadian Eastern grey squirrels: squirrelpox virus surveillance and why nothing matters.

    McInnes, C., Shuttleworth, C., Larsen, K. W., Everest, D. J., Bruemmer, C., Carroll, B., Romeo, C., Sainsbury, A. W., Crawshaw, G., Dubois, S., Gillis, L., Gilray, J. & Percival, A., 3 Aug 2020, In: Hystrix-Italian Journal of Mammalogy. 31, p. 95-98

    Research output: Contribution to journal › Article › peer-review

  31. Published

    Reintroductions and translocations of red squirrels within Europe

    Lawton, C., Waters, C. & Shuttleworth, C., 2015, Red squirrels: Ecology, Conservation & Management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Hayward, M. W. (eds.). European Squirrel Initiative, p. 193-210

    Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review

  32. Published

    Ranging behaviour, density and social structure in grey squirrels

    Lawton, C., Shuttleworth, C. & Kenward, R. E., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (eds.). European Squirrel Initiative, p. 133-152

    Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review

  33. Published

    Are novel ecosystems the only novelty of rewilding?

    Klop-Toker, K., Clulow, S., Shuttleworth, C. & Hayward, M., 1 Nov 2020, In: Restoration Ecology. 28, p. 1318-1320

    Research output: Contribution to journal › Article › peer-review

  34. Published

    Mathematical models of grey squirrel invasion: a case study on Anglesey

    Jones, H. E., White, A., Lurz, P. W. W., Boots, M. & Shuttleworth, C., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (eds.). European Squirrel Initiative, p. 235-255

    Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review

  35. Published

    Mathematical models for invasive species management: grey squirrel control on Anglesey

    Jones, H., White, A., Lurz, P. W. W. & Shuttleworth, C., 10 Sept 2017, In: Ecological Modelling. 180, p. 274-284

    Research output: Contribution to journal › Article › peer-review

  36. Published

    By trying to be everything, rewilding risks meaning nothing

    Jachowski, J. D., Bombaci, S., Caravaggi, A., Chalmers, A., Clulow, S., Griffin, A., Hayward, M., Heurich, M., Linnell, J., Marnewick, K., Moehrenschlager, A., Montgomery, A., Shuttleworth, C., Somers, M. & Weise, F., Nov 2019, In: Science. 364, p. 6438

    Research output: Contribution to journal › Letter

  37. Published

    First report of dermatophilosis in wild European red squirrels (Sciurus vulgaris)

    Holmes, P., Everest, D. J., Spiro, S., Wessels, M. & Shuttleworth, C., 30 Sept 2019, In: Veterinary Record Case Reports. 7, 3, p. 3

    Research output: Contribution to journal › Article › peer-review

  38. Published

    Repeated squirrelpox infection in Welsh red squirrels

    Holmes, J. P., Everest, D. & Shuttleworth, C., 3 Feb 2023, In: Veterinary Record. 192, 3, p. 128-129

    Research output: Contribution to journal › Letter

  39. Published

    Deconstructing compassionate conservation

    Hayward, M. W., Callen, A., Allen, B. L., Ballard, G., Broekhuis, F., Bugir, C., Clarke, R. H., Clulow, J., Clulow, S., Daltry, J. C., Davies-Mostert, H. T., Fleming, P. J. S., Griffin, A. S., Howell, L. G., Kerley, G. I. H., Klop-Toker, K., Legge, S., Major, T., Meyer, N., Montgomery, R. A., Moseby, K., Parker, D. M., Périquet, S., Read, J., Scanlon, R., Seeto, R., Shuttleworth, C., Somers, M. J., Tamessar, C. T., Tuft, K., Upton, R., Valenzuela-Molina, M., Wayne, A., Witt, R. R. & Wuster, W., Aug 2019, In: Conservation Biology. 33, 4, p. 760-768

    Research output: Contribution to journal › Article › peer-review

  40. Published

    Reintroducing rewilding to restoration – Rejecting the search for novelty

    Hayward, M. W., Scanlon, R. J., Callen, A., Howell, L. G., Klop-Toker, K. L., Di Blanco, Y., Balkenhol, N., Bugir, C. K., Campbell, L., Caravaggi, A., Chalmers, A. C., Clulow, J., Clulow, S., Cross, P., Gould, J. A., Griffin, A. S., Heurich, M., Howe, B. K., Jachowski, D. S., Jhala, Y. V., Krishnamurthy, R., Kowalczyk, R., Lenga, D. J., Linnell, J. D. C., Marnewick, K. A., Moehrenschlager, A., Montgomery, R. A., Osipova, L., Peneaux, C., Rodger, J. C., Sales, L. P., Seeto, R. G. Y., Shuttleworth, C. M., Somers, M. J., Tamessar, C. T., Upton, R. M. O. & Weise, F. J., 1 May 2019, In: Biological Conservation. 233, p. 255-259 5 p.

    Research output: Contribution to journal › Article › peer-review

  41. Published

    The search for novelty continues for rewilding

    Hayward, M., Jachowski, J. D., Bugir, C. J., Clulow, J., Krishnamurthy, R., Griffin, A. S., Chalmers, A. C., Linnell, J. D. C., Montgomery, R. A., Somers, M. J., Kowalczyk, R., Heurich, M., Caravaggi, A., Marnewick, K. A., Di Blanco, Y., Shuttleworth, C., Callen, C., Weise, F., Scanlon, R., Moehrenschlager, A., Howell, L. & Upton, R. M. O., 1 Aug 2019, In: Biological Conservation. 236, p. 584-585

    Research output: Contribution to journal › Article › peer-review

  42. Published

    Rewilding needs to clarify the role of management of invasive species

    Hayward, M., Jachowski, D. S., Shuttleworth, C., Linnell, J. D. C., Allen, B. L., Griffen, A. S., Montgomery, R. A., Caravaggi, A., Weise, F. J., Moehrenschlager, A., Somers, M. J., Clulow, S., Heurich, M. & Marnewick, K., 8 May 2019, In: Science. 364, p. 6438

    Research output: Contribution to journal › Letter

  43. Published

    Experiences of grey squirrel management in an upland conifer forest

    Halliwell, E. C., Shuttleworth, C., Cartmel, S., Lloyd, I. & Jenkins, R., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (eds.). European Squirrel Initiative, p. 453-474

    Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review

  44. Published

    Striving for success: an evaluation of local action to conserve red squirrels (Sciurus vulgaris) in Wales

    Halliwell, E. C., Shuttleworth, C., Wilberforce, L., Denman, H., Lloyd, I. & Cartmel, S., 2015, Red squirrels: Ecology, Conservation & Management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Hayward, M. W. (eds.). European Squirrel Initiative, p. 175-192

    Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review

  45. Published

    Significant differences in the caecal bacterial microbiota of red and grey squirrels in Britain

    Hall, L., Nichols, C., Leng, J., Martelli, F., Shuttleworth, C. & La Ragione, R., 14 Feb 2024, In: Journal of Medical Microbiology. 73, 2

    Research output: Contribution to journal › Article › peer-review

  46. Published

    Ecosystem impacts of an alien invader in Europe, the grey squirrel Sciurus carolinensis

    Gurnell, J., Lurz, P. W. W. & Shuttleworth, C., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (eds.). European Squirrel Initiative, p. 307-329

    Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review

  47. Published

    British red squirrel strongholds: challenges for conservation

    Gurnell, J., Blackett, T., Butler, H., Lurz, P. W. W., Magris, L. & Shuttleworth, C., 2015, Red squirrels: Ecology, Conservation & Management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Hayward, M. W. (eds.). European Squirrel Initiative, p. 211-232

    Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review

  48. Published

    Controlling grey squirrels in forests and woodlands in the UK

    Gill, R., Ferryman, M., Shuttleworth, C., Lurz, P. W. W., Mill, A., Robertson, P. A. & Dutton, C., 2019, Forest Research. 16 p.

    Research output: Book/Report › Book › peer-review

  49. Published

    PCR detection of adenovirus in grey squirrels on Anglesey

    Everest, D. J., Grierson, S. S., Stidworthy, M. & Shuttleworth, C., 31 Oct 2009, In: Veterinary Record. 165, p. 482

    Research output: Contribution to journal › Article › peer-review

  50. Published

    Adenovirus-associated deaths in red squirrels on Anglesey

    Everest, D. J., Shuttleworth, C. & Stidworthy, M. F., 31 Oct 2008, In: The Veterinary record. 163, 14, p. 430

    Research output: Contribution to journal › Article › peer-review

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