Dr Craig Shuttleworth
Honorary Visiting Research Fellow

Affiliations
Links
- https://twitter.com/Welshsquirrels
Personal account - https://twitter.com/RedSquirrelsinW
Welsh red squirrel conservation - https://www.facebook.com/groups/143610382316300
Welsh red squirrel conservation - https://pinemarten.net/
The Gwynedd pine marten project - https://www.linkedin.com/in/craig-shuttleworth-6062a1253/
Personal page
Contact info
Mobile 07966150847
Email c.shuttleworth@bangor.ac.uk
- 2025
- Published
Mitochondrial DNA Analysis of Red Squirrels (Sciurus vulgaris) in North Wales
O’Meara, D., Shuttleworth, C., Clews-Roberts, R., Everest, D. & Synnott, R., 1 Mar 2025, 14 p. (Evidence Report Series; no. 791)Research output: Book/Report › Commissioned report
- 2024
- Published
Developing a method for assessing the relative abundance of red squirrels in low-density populations
Shannon, G., Barton, O., Valle, S. & Shuttleworth, C., 1 Nov 2024, 34 p.Research output: Book/Report › Commissioned report
- Published
Gut bacteria may explain why grey squirrels outcompete reds – new research
Shuttleworth, C., 20 Feb 2024, The Conversation.Research output: Contribution to specialist publication › Article
- Published
Significant differences in the caecal bacterial microbiota of red and grey squirrels in Britain
Hall, L., Nichols, C., Leng, J., Martelli, F., Shuttleworth, C. & La Ragione, R., 14 Feb 2024, In: Journal of Medical Microbiology. 73, 2Research output: Contribution to journal › Article › peer-review
- 2023
- Published
Adenovirus remains widespread in squirrels on Anglesey
Shuttleworth, C., Everest, D. & McQueen, A., 16 Dec 2023, In: Veterinary Record. 193, 12, p. 490-491 2 p.Research output: Contribution to journal › Letter
- Published
Capturing red squirrels (Sciurus vulgaris) on camera: A cost-effective approach for monitoring relative abundance and habitat preference
Shannon, G., Valle, S. & Shuttleworth, C., 3 Oct 2023, In: Ecology and Evolution. 13, 10, e10536.Research output: Contribution to journal › Article › peer-review
- Published
Invasive genetic rescue: dispersal following repeated culling reinforces the genetic diversity of an invasive mammal
Synnott, R., Shuttleworth, C., Everest, D., Stevenson-Holt, C., O'Reilly, C., McDevitt, A. & O.Meara, D., Oct 2023, In: Biological Invasions. 25, p. 3323-3339Research output: Contribution to journal › Article › peer-review
- Published
Opportunistic viral surveillance confirms the ongoing disease threat grey squirrels pose to sympatric red squirrel populations in the UK
Everest, D., Green, C., Dastjerdi, A., Davies, H., Cripps, R., McKinney, C., Podgornik, G., Stinson, M., O'Hare, S., Bonnie, S., Mill, A., Van der wall, Z., Robinson, N., Trotter, S. & Shuttleworth, C., 20 May 2023, In: Veterinary Record. 192, 10, e2834.Research output: Contribution to journal › Article › peer-review
- Published
Repeated squirrelpox infection in Welsh red squirrels
Holmes, J. P., Everest, D. & Shuttleworth, C., 3 Feb 2023, In: Veterinary Record. 192, 3, p. 128-129Research output: Contribution to journal › Letter
- Published
Indirect effects of pine marten recovery result in benefits to native prey through suppression of an invasive species and a shared pathogen
Slade, A., White, A., Lurz, P. W. W., Shuttleworth, C., Tosh, D. & Twining, J., 1 Feb 2023, In: Ecological Modelling. 476, 110216.Research output: Contribution to journal › Article › peer-review
- Published
Restoring our lost wildlife: a review of the ecological, social and economic factors in the potential reintroduction of six mammal species to Cornwall
Cooper, P., Brazier, R., Puttock, A., Graham, H., Auster, R., Campbell-Palmer, R., Shuttleworth, C. & Pizzi, R., 2023, 258 p.Research output: Book/Report › Commissioned report
- 2022
- Published
Squirrelpox outbreak detected in north Wales – without a vaccine, the disease will keep decimating red squirrels
Shuttleworth, C., 21 Dec 2022, The Conversation.Research output: Other contribution
- Published
A temporal refuge from predation can change the outcome of prey species competition
Slade, A., White, A., Lurz, P. W. W., Shuttleworth, C. & Lambin, X., 1 Sept 2022, In: Oikos. Oikos e08565Research output: Contribution to journal › Article › peer-review
- Published
Non-invasive genetic studies and virus epidemiology
Everest, D. J., Dastjerdi, A., Davies, H., Shuttleworth, C., Ogden, R., Butler, H. & Sapsford, B., 30 Mar 2022, In: Veterinary Record. 190, p. 204-205Research output: Contribution to journal › Article
- Published
An Opportunistic Assessment of the Impact of Squirrelpox Disease Outbreaks upon a Red Squirrel Population Sympatric with Grey Squirrels in Wales
Shuttleworth, C., Everest, D., Holmes, P., Bell, S. & Cripps, R., 1 Jan 2022, In: Animals. 12, 1Research output: Contribution to journal › Article › peer-review
- 2021
- Published
Adenovirus infection in red squirrels
Shuttleworth, C., Everest, D. J., Stidworthy, M., Halliwell, E. C., Clews-Roberts, R., Paine, V., Bamber, C. & Mason, J., 31 Dec 2021, In: The Veterinary record. 189, p. 444-445Research output: Contribution to journal › Article
- Published
Disease monitoring and surveillance: case studies in the applied conservation of fragmented red squirrel (Sciurus vulgaris) populations in England and Wales
Everest, D. J., Floyd, T., Holmes, P., Man, C., Dunnett, C., Locke, R., Savage, L., Sutcliffe, S., Sapsford, B., Shuttleworth, C. & Duff, P., 1 Dec 2021, In: Mammalian Biology. 101, p. 1079-1088Research output: Contribution to journal › Article › peer-review
- Published
Sympatric cases of squirrelpox in red and grey squirrels
Shuttleworth, C., Everest, D., Wood, C. & Holmes, J. P., 7 May 2021, In: The Veterinary record. 188, 9, p. 358Research output: Contribution to journal › Article › peer-review
- Published
Recalibrating Risk: Implications of squirrelpox virus for successful red squirrel translocations within mainland UK
Shuttleworth, C., Brady, D., Cross, P., Gardner, L., Greenwood, A., Jackson, N., McKinney, C., Robinson, N., Trotter, S., Valle, S., Wood, K. & Hayward, M., Feb 2021, In: Conservation Science and Practice. 3, 2, e321.Research output: Contribution to journal › Article › peer-review
- Published
A practical guide to early surveillance and monitoring
Robinson, N. J., Everest, D. J., Harry, P. & Shuttleworth, C., 2021, Invasive Alien Species Colonisation Prevention: Your guide to early detection and rapid response . Robinson, N. J. & Shuttleworth, C. M. (eds.). Royal Society of Wildlife Trusts, p. 163-202Research output: Chapter in Book/Report/Conference proceeding › Chapter
- Published
Magical Mammals: Red squirrels in North West Wales
Shuttleworth, C., 2021, Saving the red squirrel: landscape recovery. Shuttleworth, C. M., Lurz, P. W. W. & Robinson, N. J. (eds.). Red Squirrel Survival Trust, p. 269-278Research output: Chapter in Book/Report/Conference proceeding › Chapter
- Published
Monitoring escaped captive bred red squirrels in the UK
Shuttleworth, C., 2021, Saving the red squirrel: landscape recovery. Shuttleworth, C. M., Lurz, P. W. W. & Robinson, N. J. (eds.). Red Squirrel Survival Trust, p. 71-78Research output: Chapter in Book/Report/Conference proceeding › Chapter
- Published
Preventing grey squirrel establishment on Anglesey
Shuttleworth, C., 2021, Invasive Alien Species Colonisation Prevention: Your guide to early detection and rapid response . Robinson, N. J. & Shuttleworth, C. M. (eds.). Royal Society of Wildlife Trusts, p. 55-69Research output: Chapter in Book/Report/Conference proceeding › Chapter
- Published
Red Squirrel Conservation in Clocaenog Forest: Review and Recommendations
Shuttleworth, C., 2021, 56 p.Research output: Book/Report › Commissioned report
- Published
Saving the red squirrel: landscape scale recovery
Shuttleworth, C., Lurz, P. W. W. & Robinson, N., 2021, London: Red Squirrel Survival Trust. 306 p.Research output: Book/Report › Book › peer-review
- 2020
- Published
Reinstating trophic cascades as an applied conservation tool to protect forest ecosystems from invasive grey squirrels (Sciurus carolinensis)
Bamber, J. A., Shuttleworth, C. M., Hayward, M. W. & Everest, D. J., 31 Dec 2020, In: Food Webs. 25, e00164.Research output: Contribution to journal › Article › peer-review
- Published
Are novel ecosystems the only novelty of rewilding?
Klop-Toker, K., Clulow, S., Shuttleworth, C. & Hayward, M., 1 Nov 2020, In: Restoration Ecology. 28, p. 1318-1320Research output: Contribution to journal › Article › peer-review
- Published
Ethical dilemmas when using citizen science for early detection of invasive tree pests and diseases
Pocock, M. J. O., Marzano, M., Bullas-Appleton, E., Dyke, A., de Groot, M., Shuttleworth, C. & White, R., 1 Nov 2020, In: Management of Biological Invasions. 11, p. 720-732Research output: Contribution to journal › Article › peer-review
- Published
Evolving grey squirrel management techniques in Europe
Shuttleworth, C., Robinson, N. J., Halliwell, E. C., Clews-Roberts, R., Peek, H., Podgornik, G., Stinson, M., Rice, S., Finlay, C., McKinney, C., Everest, D. J. & Larsen, K. W., 21 Oct 2020, In: Management of Biological Invasions. 11, 4, p. 747-761Research output: Contribution to journal › Article › peer-review
- Published
Do differing levels of boldness influence the success of translocation? A pilot study on red squirrels (Sciurus vulgaris)
Bamber, J. A., Shuttleworth, C. & Hayward, M., 26 Sept 2020, In: Animals. 10, 1748.Research output: Contribution to journal › Article › peer-review
- Published
Challenges and solutions in early detection, rapid response and communication about potential invasive alien species in forests
de Groot, M., Bullas-Appleton, E., Csóka, G., Csiszár, A., Faccoli, A., Gervasini, A., Kirichenko, N., Shuttleworth, C., Kus Veenvliet, J., O'Hanlon, R., Korda, M., Marinsek, A., Robinson, N., Sweeney, J., Tricarico, E., Verbrugge, L., Williams, D. & Zidar, S., 14 Aug 2020, In: Management of Biological Invasions. 11, p. 637-660Research output: Contribution to journal › Article › peer-review
- Published
Introduced Canadian Eastern grey squirrels: squirrelpox virus surveillance and why nothing matters.
McInnes, C., Shuttleworth, C., Larsen, K. W., Everest, D. J., Bruemmer, C., Carroll, B., Romeo, C., Sainsbury, A. W., Crawshaw, G., Dubois, S., Gillis, L., Gilray, J. & Percival, A., 3 Aug 2020, In: Hystrix-Italian Journal of Mammalogy. 31, p. 95-98Research output: Contribution to journal › Article › peer-review
- Published
Response to comments on "Compassionate Conservation deserves a morally serious rather than dismissive response - reply to Callen et al., 2020"
Callen, A., Hayward, M. W., Klop-Toker, K., Allen, B. L., Ballard, G., Beranek, C. T., Broekhuis, F., Bugir, C. K., Clarke, R. H., Clulow, J., Clulow, S., Daltry, J. C., Davies-Mostert, H. T., Di Blanco, Y. E., Dixon, V., Fleming, P. J. S., Howell, L. G., Kerley, G. I. H., Legge, S. M., Lenga, D. J., Major, T., Montgomery, R. A., Moseby, K., Meyer, N., Parker, D. M., Periquet, S., Read, J., Scanlon, R. J., Shuttleworth, C., Tamessar, C. T., Taylor, W. A., Tuft, K., Upton, R. M. O., Valenzuela, M., Witt, R. R. & Wuster, W., 30 Apr 2020, In: Biological Conservation. 244Research output: Contribution to journal › Letter › peer-review
- Published
Envisioning the future with 'compassionate conservation': An ominous projection for native wildlife and biodiversity
Callen, A., Hayward, M. W., Klop-Toker, K., Allen, B. L., Ballard, G., Beranek, C. T., Broekhuis, F., Bugir, C. K., Clarke, R. H., Clulow, J., Clulow, S., Daltry, J. C., Davies-Mostert, H. T., Di Blanco, Y. E., Dixon, V., Fleming, P. J. S., Howell, L. G., Kerley, G. I. H., Legge, S. M., Lenga, D. J., Major, T., Montgomery, R. A., Moseby, K., Meyer, N., Parker, D. M., Periquet, S., Read, J., Scanlon, R. J., Shuttleworth, C., Tamessar, C. T., Taylor, W. A., Tuft, K., Upton, R. M. O., Valenzuela, M., Witt, R. R. & Wuster, W., Jan 2020, In: Biological Conservation. 241, 108365.Research output: Contribution to journal › Article › peer-review
- Published
Invasive alien species colonisation prevention: Your guide to early detection and rapid response
Robinson, N. & Shuttleworth, C., 2020, Newark: Royal Society of Wildlife Trusts. 204 p.Research output: Book/Report › Book › peer-review
- 2019
- Published
Pine marten scat holds few clues for squirrel disease
Shuttleworth, C., Everest, D. J., Dastjerdi, A., Birks, J. & Hayward, M., 7 Dec 2019, In: Veterinary Record. 185, p. 698Research output: Contribution to journal › Article
- Published
By trying to be everything, rewilding risks meaning nothing
Jachowski, J. D., Bombaci, S., Caravaggi, A., Chalmers, A., Clulow, S., Griffin, A., Hayward, M., Heurich, M., Linnell, J., Marnewick, K., Moehrenschlager, A., Montgomery, A., Shuttleworth, C., Somers, M. & Weise, F., Nov 2019, In: Science. 364, p. 6438Research output: Contribution to journal › Letter
- Published
First report of dermatophilosis in wild European red squirrels (Sciurus vulgaris)
Holmes, P., Everest, D. J., Spiro, S., Wessels, M. & Shuttleworth, C., 30 Sept 2019, In: Veterinary Record Case Reports. 7, 3, p. 3Research output: Contribution to journal › Article › peer-review
- Published
The search for novelty continues for rewilding
Hayward, M., Jachowski, J. D., Bugir, C. J., Clulow, J., Krishnamurthy, R., Griffin, A. S., Chalmers, A. C., Linnell, J. D. C., Montgomery, R. A., Somers, M. J., Kowalczyk, R., Heurich, M., Caravaggi, A., Marnewick, K. A., Di Blanco, Y., Shuttleworth, C., Callen, C., Weise, F., Scanlon, R., Moehrenschlager, A., Howell, L. & Upton, R. M. O., 1 Aug 2019, In: Biological Conservation. 236, p. 584-585Research output: Contribution to journal › Article › peer-review
- Published
Deconstructing compassionate conservation
Hayward, M. W., Callen, A., Allen, B. L., Ballard, G., Broekhuis, F., Bugir, C., Clarke, R. H., Clulow, J., Clulow, S., Daltry, J. C., Davies-Mostert, H. T., Fleming, P. J. S., Griffin, A. S., Howell, L. G., Kerley, G. I. H., Klop-Toker, K., Legge, S., Major, T., Meyer, N., Montgomery, R. A., Moseby, K., Parker, D. M., Périquet, S., Read, J., Scanlon, R., Seeto, R., Shuttleworth, C., Somers, M. J., Tamessar, C. T., Tuft, K., Upton, R., Valenzuela-Molina, M., Wayne, A., Witt, R. R. & Wuster, W., Aug 2019, In: Conservation Biology. 33, 4, p. 760-768Research output: Contribution to journal › Article › peer-review
- Published
Assessing a potential non-invasive method for viral diagnostic purposes in European squirrels
Everest, D. J., Tolhurst-Cherriman, D. A. R., Davies, H., Dastjerdi, A., Ashton, A., Blackett, T., Meredith, A. L., Milne, E., Mill, A. & Shuttleworth, C., 4 Jun 2019, In: Hystrix-Italian Journal of Mammalogy. 30, p. 44-50Research output: Contribution to journal › Article › peer-review
- Published
Rewilding needs to clarify the role of management of invasive species
Hayward, M., Jachowski, D. S., Shuttleworth, C., Linnell, J. D. C., Allen, B. L., Griffen, A. S., Montgomery, R. A., Caravaggi, A., Weise, F. J., Moehrenschlager, A., Somers, M. J., Clulow, S., Heurich, M. & Marnewick, K., 8 May 2019, In: Science. 364, p. 6438Research output: Contribution to journal › Letter
- Published
Reintroducing rewilding to restoration – Rejecting the search for novelty
Hayward, M. W., Scanlon, R. J., Callen, A., Howell, L. G., Klop-Toker, K. L., Di Blanco, Y., Balkenhol, N., Bugir, C. K., Campbell, L., Caravaggi, A., Chalmers, A. C., Clulow, J., Clulow, S., Cross, P., Gould, J. A., Griffin, A. S., Heurich, M., Howe, B. K., Jachowski, D. S., Jhala, Y. V., Krishnamurthy, R., Kowalczyk, R., Lenga, D. J., Linnell, J. D. C., Marnewick, K. A., Moehrenschlager, A., Montgomery, R. A., Osipova, L., Peneaux, C., Rodger, J. C., Sales, L. P., Seeto, R. G. Y., Shuttleworth, C. M., Somers, M. J., Tamessar, C. T., Upton, R. M. O. & Weise, F. J., 1 May 2019, In: Biological Conservation. 233, p. 255-259 5 p.Research output: Contribution to journal › Article › peer-review
- Published
Detecting viral infection in red squirrels
Shuttleworth, C., Everest, D. J., Halliwell, E. C., Hulme, B., Wilberforce, L. & Clews-Roberts, R., 30 Apr 2019, In: Veterinary Record. 184, p. 507Research output: Contribution to journal › Article
- Published
Releasing grey squirrels into the wild
Shuttleworth, C., Bertolino, S., Gill, R., Gurnell, J., Hayward, M., Kenward, R. E., Lawton, C., Lurz, P. W. W., McInnes, C. J., Mill, A., Trotter, S. & Wauters, L. A., 31 Mar 2019, In: Veterinary Record. 184, p. 389-390Research output: Contribution to journal › Article
- Published
Leprosy in red squirrels in the UK
Schilling, A.-K., Del-Pozo, J., Lurz, P. W. W., Stevenson, K., Avanzi, C., Shuttleworth, C., Cole, S. T. & Meredith, A. L., 30 Mar 2019, In: Veterinary Record. 184, p. 416Research output: Contribution to journal › Article
- Published
British red squirrels remain the only known wild rodent host for leprosy bacilli
Schilling, A., Avanzi, C., Ulrich, R. G., Busso, P., Pisanu, B., Ferrari, N., Romeo, C., Mazzamuto, M. V., McLuckie, J., Shuttleworth, C., Del-Pozo, J., Lurz, P. W. W., Escalante-Fuentes, W. G., Ocampo-Candiani, J., Vera-Cabrera, L., Stevenson, K., Chapuis, J.-L., Meredith, A. L. & Cole, S. T., 1 Feb 2019, In: Frontiers in Veterinary Science. 6, p. 8Research output: Contribution to journal › Article › peer-review
- Published
Disease, invasions and conservation: no evidence of squirrelpox virus in grey squirrels introduced to Italy
Romeo, C., McInnes, C., Dale, T., Shuttleworth, C., Bertolino, S., Wauters, L. A. & Ferrari, N., 1 Feb 2019, In: Animal Conservation. 22, 1, p. 14-23Research output: Contribution to journal › Article › peer-review
- Published
Controlling grey squirrels in forests and woodlands in the UK
Gill, R., Ferryman, M., Shuttleworth, C., Lurz, P. W. W., Mill, A., Robertson, P. A. & Dutton, C., 2019, Forest Research. 16 p.Research output: Book/Report › Book › peer-review
- Published
Red squirrel bark-stripping of hornbeam in North Wales
Shuttleworth, C. & Gill, R., 2019, In: Quarterly Journal of Forestry. 113, 4, p. 275-277Research output: Contribution to journal › Article › peer-review