Research Portal

Dr Craig Shuttleworth

Honorary Visiting Research Fellow

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ORCID: 0000-0003-0076-6789

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Mobile 07966150847

Email c.shuttleworth@bangor.ac.uk

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  1. Article › Research › Peer-reviewed
  2. Published

    Reducing negative impacts on non-target species during grey squirrel (Sciurus carolinensis) control activities

    Shuttleworth, C., Mill, A. C., Van der Wall, Z., Bertolino, S. & Everest, D. J., 1 Jan 2018, In: Quarterly Journal of Forestry. 112, 1, p. 21-29

    Research output: Contribution to journal › Article › peer-review

  3. Published

    Reinstating trophic cascades as an applied conservation tool to protect forest ecosystems from invasive grey squirrels (Sciurus carolinensis)

    Bamber, J. A., Shuttleworth, C. M., Hayward, M. W. & Everest, D. J., 31 Dec 2020, In: Food Webs. 25, e00164.

    Research output: Contribution to journal › Article › peer-review

  4. Published

    Reintroducing rewilding to restoration – Rejecting the search for novelty

    Hayward, M. W., Scanlon, R. J., Callen, A., Howell, L. G., Klop-Toker, K. L., Di Blanco, Y., Balkenhol, N., Bugir, C. K., Campbell, L., Caravaggi, A., Chalmers, A. C., Clulow, J., Clulow, S., Cross, P., Gould, J. A., Griffin, A. S., Heurich, M., Howe, B. K., Jachowski, D. S., Jhala, Y. V., Krishnamurthy, R., Kowalczyk, R., Lenga, D. J., Linnell, J. D. C., Marnewick, K. A., Moehrenschlager, A., Montgomery, R. A., Osipova, L., Peneaux, C., Rodger, J. C., Sales, L. P., Seeto, R. G. Y., Shuttleworth, C. M., Somers, M. J., Tamessar, C. T., Upton, R. M. O. & Weise, F. J., 1 May 2019, In: Biological Conservation. 233, p. 255-259 5 p.

    Research output: Contribution to journal › Article › peer-review

  5. Retrospective detection by negative contrast electron microscopy of faecal viral particles in wild red squirrels (Sciurus vulgaris) with suspected enteropathy in Great Britain

    Everest, D. J., Stidworthy, M. F., Milne, E. M., Meredith, A. L., Chantery, J., Shuttleworth, C., Blackett, T., Butler, H., Wilkinson, M. & Sainsbury, A. W., 31 Dec 2010, In: Veterinary Record. 167, p. 1007-1010

    Research output: Contribution to journal › Article › peer-review

  6. Published

    Significant differences in the caecal bacterial microbiota of red and grey squirrels in Britain

    Hall, L., Nichols, C., Leng, J., Martelli, F., Shuttleworth, C. & La Ragione, R., 14 Feb 2024, In: Journal of Medical Microbiology. 73, 2

    Research output: Contribution to journal › Article › peer-review

  7. Published

    Sympatric cases of squirrelpox in red and grey squirrels

    Shuttleworth, C., Everest, D., Wood, C. & Holmes, J. P., 7 May 2021, In: The Veterinary record. 188, 9, p. 358

    Research output: Contribution to journal › Article › peer-review

  8. Published

    The foraging behaviour and diet of red squirrels (Sciurus vulgaris) receiving supplemental feeding

    Shuttleworth, C., 1 Sept 2000, In: Wildlife Biology. 6, p. 149-156

    Research output: Contribution to journal › Article › peer-review

  9. Published

    The implications of significant adenovirus infection in UK captive red squirrel (Sciurus vulgaris) collections: How histological screening can aid applied conservation management

    Everest, D. J., Shuttleworth, C., Grierson, S. S., Dastjerdi, A., M.F., S., Duff, J. P., Higgins, R. J., Mill, A. C. & Chantrey, J., 1 Jan 2018, In: Mammalian Biology. 88, p. 123-129

    Research output: Contribution to journal › Article › peer-review

  10. Published

    The large-scale removal of mammalian invasive alien species in Northern Europe

    Robertson, P. A., Adriaens, T., Lambin, X., Mill, A., Roy, S., Shuttleworth, C. & Sutton-Croft, M., 1 Feb 2017, In: Pest Management Science. 73, p. 273-279

    Research output: Contribution to journal › Article › peer-review

  11. Published

    The search for novelty continues for rewilding

    Hayward, M., Jachowski, J. D., Bugir, C. J., Clulow, J., Krishnamurthy, R., Griffin, A. S., Chalmers, A. C., Linnell, J. D. C., Montgomery, R. A., Somers, M. J., Kowalczyk, R., Heurich, M., Caravaggi, A., Marnewick, K. A., Di Blanco, Y., Shuttleworth, C., Callen, C., Weise, F., Scanlon, R., Moehrenschlager, A., Howell, L. & Upton, R. M. O., 1 Aug 2019, In: Biological Conservation. 236, p. 584-585

    Research output: Contribution to journal › Article › peer-review

  12. Published

    The use of nest boxes by the red squirrel (Sciurus vulgaris) in conifer plantations

    Shuttleworth, C., 2001, In: Quarterly Journal of Forestry. 95, p. 225-228

    Research output: Contribution to journal › Article › peer-review

  13. Published

    The use of nest boxes by the red squirrel Sciurus vulgaris in a coniferous habitat

    Shuttleworth, C., 1 Mar 1999, In: Mammal Review. 29, p. 61-66

    Research output: Contribution to journal › Article › peer-review

  14. Published

    Traffic related mortality in a red squirrel (Sciurus vulgaris) population receiving supplemental feeding

    Shuttleworth, C., 1 Jun 2001, In: Urban Ecosystems. 2, p. 109-118

    Research output: Contribution to journal › Article › peer-review

  15. Letter › Research › Not peer-reviewed
  16. Published

    Adenovirus remains widespread in squirrels on Anglesey

    Shuttleworth, C., Everest, D. & McQueen, A., 16 Dec 2023, In: Veterinary Record. 193, 12, p. 490-491 2 p.

    Research output: Contribution to journal › Letter

  17. Published

    By trying to be everything, rewilding risks meaning nothing

    Jachowski, J. D., Bombaci, S., Caravaggi, A., Chalmers, A., Clulow, S., Griffin, A., Hayward, M., Heurich, M., Linnell, J., Marnewick, K., Moehrenschlager, A., Montgomery, A., Shuttleworth, C., Somers, M. & Weise, F., Nov 2019, In: Science. 364, p. 6438

    Research output: Contribution to journal › Letter

  18. Published

    Repeated squirrelpox infection in Welsh red squirrels

    Holmes, J. P., Everest, D. & Shuttleworth, C., 3 Feb 2023, In: Veterinary Record. 192, 3, p. 128-129

    Research output: Contribution to journal › Letter

  19. Published

    Rewilding needs to clarify the role of management of invasive species

    Hayward, M., Jachowski, D. S., Shuttleworth, C., Linnell, J. D. C., Allen, B. L., Griffen, A. S., Montgomery, R. A., Caravaggi, A., Weise, F. J., Moehrenschlager, A., Somers, M. J., Clulow, S., Heurich, M. & Marnewick, K., 8 May 2019, In: Science. 364, p. 6438

    Research output: Contribution to journal › Letter

  20. Letter › Research › Peer-reviewed
  21. Published

    Response to comments on "Compassionate Conservation deserves a morally serious rather than dismissive response - reply to Callen et al., 2020"

    Callen, A., Hayward, M. W., Klop-Toker, K., Allen, B. L., Ballard, G., Beranek, C. T., Broekhuis, F., Bugir, C. K., Clarke, R. H., Clulow, J., Clulow, S., Daltry, J. C., Davies-Mostert, H. T., Di Blanco, Y. E., Dixon, V., Fleming, P. J. S., Howell, L. G., Kerley, G. I. H., Legge, S. M., Lenga, D. J., Major, T., Montgomery, R. A., Moseby, K., Meyer, N., Parker, D. M., Periquet, S., Read, J., Scanlon, R. J., Shuttleworth, C., Tamessar, C. T., Taylor, W. A., Tuft, K., Upton, R. M. O., Valenzuela, M., Witt, R. R. & Wuster, W., 30 Apr 2020, In: Biological Conservation. 244

    Research output: Contribution to journal › Letter › peer-review

  22. Chapter › Research › Not peer-reviewed
  23. Published

    A practical guide to early surveillance and monitoring

    Robinson, N. J., Everest, D. J., Harry, P. & Shuttleworth, C., 2021, Invasive Alien Species Colonisation Prevention: Your guide to early detection and rapid response . Robinson, N. J. & Shuttleworth, C. M. (eds.). Royal Society of Wildlife Trusts, p. 163-202

    Research output: Chapter in Book/Report/Conference proceeding › Chapter

  24. Published

    Against a grey tide: What future for the red squirrel?

    Shuttleworth, C., 2005, Journal of Practical Ecology & Conservation (Special Series No 4): Living with the Aliens? - Ecology, History, Culture and Management if exotic plants and animals: issues for nature conservation . Rotherham, L. D. (ed.). Vol. 4. p. 87-88

    Research output: Chapter in Book/Report/Conference proceeding › Chapter

  25. Published

    Developing red squirrel re-introduction techniques for use during regional grey squirrel eradication programmes in Europe

    Shuttleworth, C., Jackson, N. & Kenward, R. E., 2015, 2015 Global Reintroduction Perspectives. Soorae, P. (ed.). IUCN/SSC Re-introduction Specialist Group, p. 67-74

    Research output: Chapter in Book/Report/Conference proceeding › Chapter

  26. Published

    Island Haven: Anglesey red squirrel conservation

    Shuttleworth, C., 2015, Shared Experience of Red Squirrel Conservation Practice . Shuttleworth, C. M., Lurz, P. W. W. & Halliwell, E. C. (eds.). European Squirrel Initiative, p. 151-158

    Research output: Chapter in Book/Report/Conference proceeding › Chapter

  27. Published

    Magical Mammals: Red squirrels in North West Wales

    Shuttleworth, C., 2021, Saving the red squirrel: landscape recovery. Shuttleworth, C. M., Lurz, P. W. W. & Robinson, N. J. (eds.). Red Squirrel Survival Trust, p. 269-278

    Research output: Chapter in Book/Report/Conference proceeding › Chapter

  28. Published

    Monitoring escaped captive bred red squirrels in the UK

    Shuttleworth, C., 2021, Saving the red squirrel: landscape recovery. Shuttleworth, C. M., Lurz, P. W. W. & Robinson, N. J. (eds.). Red Squirrel Survival Trust, p. 71-78

    Research output: Chapter in Book/Report/Conference proceeding › Chapter

  29. Published

    Painting the Town Red: Gwynedd red squirrel conservation

    Shuttleworth, C., 2015, Shared Experience of Red Squirrel Conservation Practice . Shuttleworth, C. M., Lurz, P. W. W. & Halliwell, E. C. (eds.). European Squirrel Initiative, p. 159-166

    Research output: Chapter in Book/Report/Conference proceeding › Chapter

  30. Published

    Preventing grey squirrel establishment on Anglesey

    Shuttleworth, C., 2021, Invasive Alien Species Colonisation Prevention: Your guide to early detection and rapid response . Robinson, N. J. & Shuttleworth, C. M. (eds.). Royal Society of Wildlife Trusts, p. 55-69

    Research output: Chapter in Book/Report/Conference proceeding › Chapter

  31. Published

    Re-introduction of the red squirrel into Newborough forest on the island of Anglesey, UK

    Shuttleworth, C., Kenward, R. E. & Jackson, N., 2008, Global Re-Introduction Perspectives: Re-Introduction Case-Studies from Around the Globe. Soorae, P. S. (ed.). IUCN/SSC Re-introduction Specialist Group, p. 163-166

    Research output: Chapter in Book/Report/Conference proceeding › Chapter

  32. Published

    The management of coastal sand dune woodland for red squirrels (Sciurus vulgaris L.)

    Shuttleworth, C. & Gurnell, J., 2001, Coastal Dune Management: Shared Experience of European Conservation Practice. Houston, J., Edmondson, A. S. E. & Rooney, P. J. (eds.). Liverpool University Press, p. 117-127

    Research output: Chapter in Book/Report/Conference proceeding › Chapter

  33. Chapter › Research › Peer-reviewed
  34. Published

    British red squirrel strongholds: challenges for conservation

    Gurnell, J., Blackett, T., Butler, H., Lurz, P. W. W., Magris, L. & Shuttleworth, C., 2015, Red squirrels: Ecology, Conservation & Management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Hayward, M. W. (eds.). European Squirrel Initiative, p. 211-232

    Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review

  35. Published

    Developing integrated and applied red squirrel conservation programmes: What lessons can Europe learn from a regional grey squirrel eradication programme in North Wales?

    Shuttleworth, C., Schuchert, P., Everest, D. J., McInnes, C. J., Rushton, S. P., Jackson, N. & Kenward, R. E., 2015, Red squirrels: Ecology, Conservation & Management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Hayward, M. W. (eds.). European Squirrel Initiative, p. 233-250

    Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review

  36. Published

    Ecosystem impacts of an alien invader in Europe, the grey squirrel Sciurus carolinensis

    Gurnell, J., Lurz, P. W. W. & Shuttleworth, C., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (eds.). European Squirrel Initiative, p. 307-329

    Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review

  37. Published

    Experiences of grey squirrel management in an upland conifer forest

    Halliwell, E. C., Shuttleworth, C., Cartmel, S., Lloyd, I. & Jenkins, R., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (eds.). European Squirrel Initiative, p. 453-474

    Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review

  38. Published

    Genetic evidence of the effectiveness of grey squirrel control operations: lessons from the Isle of Anglesey

    Signorile, L. & Shuttleworth, C., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (eds.). European Squirrel Initiative, p. 439-452

    Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review

  39. Published

    Grey squirrel control as a tactic in the conservation of the red squirrel: a positive outcome

    Shuttleworth, C., 2001, Advances in Vertebrate Pest Management . Feare, C. J. & Cowan, D. P. (eds.). Filander Verlag, Vol. 3. p. 151-168

    Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review

  40. Published

    Grey squirrel management in Europe - the future

    Shuttleworth, C., Lurz, P. W. W. & Gurnell, J., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (eds.). European Squirrel Initiative

    Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review

  41. Published

    Grey squirrel nesting ecology and the use of nest sites in European population management

    Shuttleworth, C., Selonen, V. & Koprowski, J. L., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (eds.). European Squirrel Initiative, p. 349-369

    Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review

  42. Published

    Identifying incursion pathways, early detection responses and management actions to prevent grey squirrel range expansion: an island case study in Wales

    Shuttleworth, C., Halliwell, E. C. & Robertson, P. A., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (eds.). European Squirrel Initiative, p. 475-494

    Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review

  43. Published

    Mathematical models of grey squirrel invasion: a case study on Anglesey

    Jones, H. E., White, A., Lurz, P. W. W., Boots, M. & Shuttleworth, C., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (eds.). European Squirrel Initiative, p. 235-255

    Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review

  44. Published

    Ranging behaviour, density and social structure in grey squirrels

    Lawton, C., Shuttleworth, C. & Kenward, R. E., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (eds.). European Squirrel Initiative, p. 133-152

    Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review

  45. Published

    Red squirrels (Sciurus vulgaris) on the Sefton Coast: The conservation value of nest boxes

    Shuttleworth, C., 1995, 2nd NPI Red Alert UK Forum for Red Squirrel Conservation. Hughes, D. G. & Tew, T. (eds.). The Federation of Zoological Gardens of Great Britain and Ireland, p. 25-32

    Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review

  46. Published

    Reintroductions and translocations of red squirrels within Europe

    Lawton, C., Waters, C. & Shuttleworth, C., 2015, Red squirrels: Ecology, Conservation & Management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Hayward, M. W. (eds.). European Squirrel Initiative, p. 193-210

    Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review

  47. Published

    Striving for success: an evaluation of local action to conserve red squirrels (Sciurus vulgaris) in Wales

    Halliwell, E. C., Shuttleworth, C., Wilberforce, L., Denman, H., Lloyd, I. & Cartmel, S., 2015, Red squirrels: Ecology, Conservation & Management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Hayward, M. W. (eds.). European Squirrel Initiative, p. 175-192

    Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review

  48. Published

    The effect of supplemental feeding on the diet, population density and reproduction of red squirrels

    Shuttleworth, C., 1997, The Conservation of Red Squirrels (Sciurus vulgaris L.). Gurnell, J. & Lurz, P. W. W. (eds.). Peoples Trust for Endangered Species, p. 13-24

    Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review

  49. Published

    The impact of adenoviruses on red squirrel (Sciurus vulgaris) conservation – with notes on other enteric viruses of potential significance

    Everest, D. J., Shuttleworth, C., Grierson, S. S., Dastjerdi, A., Stidworthy, M. F., Duff, J. P. & Sainsbury, A. W., 2015, Red squirrels: Ecology, Conservation & Management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Hayward, M. W. (eds.). European Squirrel Initiative, p. 129-146

    Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review

  50. Published

    The management of grey squirrel populations in Europe: evolving best practice

    Bertolino, S., Lurz, P. W. W., Shuttleworth, C., Martinoli, A. & Wauters, L. A., 2016, The grey squirrel: ecology and management in Europe . Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (eds.). European Squirrel Initiative, p. 495-516

    Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review

  51. Published

    The potential role of ectoparasites in the epidemiology of squirrelpox virus: a possible novel means of intervention to reduce the impact of the disease?

    Cowan, D. P., Mill, A. C., Everest, D. J., McInnes, C. J., Rushton, S. P., Shirley, M. D. F., Start, C. & Shuttleworth, C., 2016, The grey squirrel: ecology and management in Europe . Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (eds.). European Squirrel Initiative, p. 255-289

    Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review

  52. Published

    The role of spruce in red squirrel (Sciurus vulgaris) conservation in the UK

    Shuttleworth, C. & Lurz, P. W. W., 2014, Forest Ecosystems: Biodiversity, Management and Conservation. Roberts, N. C. (ed.). Nova Science Publishers, p. 63-78

    Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review

  53. Article › Research
  54. Published

    Gut bacteria may explain why grey squirrels outcompete reds – new research

    Shuttleworth, C., 20 Feb 2024, The Conversation.

    Research output: Contribution to specialist publication › Article

  55. Other contribution › Research › Not peer-reviewed
  56. Published

    Grey squirrels are bad for the British countryside – full stop

    Shuttleworth, C., 2017, The Conversation.

    Research output: Other contribution

  57. Published

    Rare woodland wildlife at risk because of 50-year-old tree felling rules

    Shuttleworth, C., 2018, The Conversation.

    Research output: Other contribution

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