Prof John Healey

Professor of Forest Sciences


Contact info

Thoday Building, room S7


Tel: 01248 383703 (from U.K.)
+44 1248 383703 (International)


Google Scholar



Research and teaching focus: Forest ecology


For more information about my work click on "Research Interests" below


Responsibilities within BU

College of Environmental Sciences and Engineering, Director of Research

Bangor lead for the Envision NERC Doctoral Training Partnership


Ecology, management and ecosystem services of forests and agroforestry systems, with particular emphasis on tropical and upland environments. Biodiversity conservation in protection and production forests; invasive woody species. Forest restoration and remediation of degraded and post-industrial land. Sustainable forest management and policy including: climate change mitigation; Reduced Emissions from Deforestation and forest Degradation; reduced impact logging; silviculture; secondary and buffer-zone forest management; community participation; monitoring. Forest ecology including: natural regeneration; resistance and resilience of forests to disturbance; succession; landscape; climate change adaptation.  Ecosystem ecology across a range of forest, agroforestry, agricultural and upland land uses: nutrient and carbon cycling; sustainable intensification.



ForestModelling: Modelling economic impact and strategies to increase resilience against tree disease outbreaks. ForestModelling aims to build a series of models to study the ways in which different management options can reduce risks and expected damages from a range of forest diseases. It is a project of the Tree Health and Plant Biosecurity Initiative.

Invasive Woody Plants in the Tropics: The Invasive Woody Plants in the Tropics research group, based at the University of Wales, Bangor, UK, has undertaken two major field studies in Jamaica and Tanzania, together with a global review of invasions resulting in a comprehensive database, an analysis of common characteristics and recommendations for management and control.

Biodiversity conservation of ancient church and monastery yards in Ethiopia

Building Nicaraguan and Costa Rican capacity in biodiversity conservation: The homepage of the Darwin Initiative project 'Building Nicaraguan and Costa Rican capacity in biodiversity conservation'.

TWIRLS: Treating Waste for Restoring Land Sustainability: A European Commission LIFE-Environment project targeting the restoration of degraded land using locally sourced waste materials.

Cefn Conwy: The CEFN Conwy project is designed to enhance the technical knowledge and environmental management skills of farmers in rural Conwy

Envision: Developing next generation leaders in environmental science: A NERC Doctoral Training Partnership between Lancaster, Bangor and Nottingham Universities, Rothamsted Research, Centre for Ecology and Hydrology, and British Geological Survey.




Amy Gresham.  Quantifying the effects of deer on woodland structure in a human altered landscape.  Co-supervised with Graeme Shannon and Markus Eichhorn

Anita Weissflog.  Assessing the role of fungus-mediated plant-soil feedbacks during secondary succession of tropical forests. Co-supervised with Lars Markesteijn and Andy Smith

Eilidh Forster. Developing environmentally sustainable forestry value chains. Co-supervised with Dave Styles and Caren Dymond

Estrella Yungas. Developing sustainable forestry value chains in Wales. Co-supervised with Dave Styles

Owain Barton.  Environmental and human drivers of fallow deer movement and habitat use in North Wales: Implications for natural resource management. Co-supervised with Graeme Shannon and Line Cordes

Charlotte Smith. A multi-scale assessment of the regrowth potential of secondary forests in the Brazilian Amazon. Co-supervised with Fernando Espirito-Santo and Jos Barlow

Helen Ford. Widening our view of the reef: the landscape ecology of disturbance and recovery on Pacific coral reefs. Co-supervised with Gareth Williams and Nick Graham

Mark Mosbery. Temperate lianes: Forest ecology and urban habitat enrichment. Co-supervised with Lars Markesteijn

Valentino Govigli. Modelling socio-ecological interactions of sacred forests in Northern Greece. Co-supervised with John Halley and Jenny Wong




Below my journal papers published since 2012 are classified into seven subject areas:

  • ·        Tropical forest management, degradation and conservation
  • ·        Tropical forest ecology and restoration
  • ·        Agroforestry
  • ·        Temperate forest, heathland and peatland ecology and conservation
  • ·        Tree disease bioeconomic modelling
  • ·        Nutrient cycling/resource use/sustainable production
  • ·        Science-policy interface

As a cross-cutting theme, the papers directly related to climate change mitigation and adaptation are indicated by C.


Tropical forest management, degradation and conservation

Bellingham, P.J., Tanner, E.V.J., Martin, P.H., Healey, J.R. & Burge, O.R. (2018). Endemic trees in a tropical biodiversity hotspot imperilled by an invasive tree. Biological Conservation 217, 47-53 (

Kleinschroth, F., Healey, J.R. Gourlet-Fleury, S., Mortier, F. & Stoica, R.S. (2017). Effects of logging on roadless space in intact forest landscapes of the Congo Basin. Conservation Biology 31, 469-480 (

Kleinschroth, F. & Healey, J.R. (2017). Impacts of logging roads on tropical forests. Biotropica 49, 620-635(

Kleinschroth, F., Healey, J.R., Sist, P., Mortier, F. & Gourlet-Fleury, S. (2016). How persistent are the impacts of logging roads on Central African forest vegetation? Journal of Applied Ecology 53, 1127-1137.  DOI: 10.1111/1365-2664.12661. C

Kleinschroth, F., Healey, J.R. & Gourlet-Fleury, S. (2016). Sparing forests in Central Africa: re-use old logging roads to avoid creating new ones. Frontiers in Ecology and the Environment 14, 9-10 (

Kleinschroth, F., Gourlet-Fleury, S., Gond, V., Sist, P. & Healey, J.R. (2016). Logging roads in tropical forests: Synthesis of literature written in French and English highlights environmental impact reduction through improved engineering.  Bois et Forêts des Tropiques 328 (2), 13-26.

Kleinschroth, F., Gourlet-Fleury, S., Sist, P., Mortier, F. & Healey, J.R. (2015). Legacy of logging roads in the Congo Basin: how persistent are the scars in forest cover? Ecosphere 6(4): 64 (

Aerts, R., Van Overtveld, K., Wassie, A., Abiyu, A., Demissew, S., Daye, D.D., Giday, K., Haile, M., TewoldeBerhan, S., Teketay, D., Teklehaimanot, Z., Binggeli, P., Deckers, S., Friis, I., Gratzer, G., Hermy, M., Sterck, F.J., Muys, B., Bongers, F. & Healey, J.R. (2016). Conservation of the Ethiopian church forests: threats, opportunities and implications for their management. Science of the Total Environment 551-552, 404-414.  DOI: 10.1016/j.scitotenv.2016.02.034

Daye, D.D. & Healey, J.R. (2015).  Impacts of land-use change on sacred forests at the landscape scale. Global Ecology and Conservation 3, 349-358 (

Desissa, D. & Helaey, J.R. (2012). Sacred groves in the Gamo Highlands of southern Ethiopia: biocultural values and threats. Langscape 2 (11), 44-47.

Morales-Barquero, L., Borrego, A., Skutsch, M., Kleinn, C. & Healey, J.R. (2015). Identification and quantification of drivers of forest degradation in tropical dry forests: a case study in Western Mexico. Land Use Policy 49, 296-309 ( C

Morales-Barquero, L., Skutsch, M., Jardel-Peláez, E.J., Ghilardi, A., Kleinn, C. & Healey, J.R. (2014). Operationalizing the definition of forest degradation for REDD+, with application to Mexico. Forests 5, 1653-1681 (doi: 10.3390/f507 1653; C

Petrokofsky, G., Sist, P., Blanc, L., Doucet, J.-L., Finegan, B., Gourlet-Fleury, S., Healey, J.R., Livoreil, B., Nasi, R., Peña-Claros, M., Putz, F.E. & Zhou, W. (2015). Comparative effectiveness of silvicultural interventions for increasing timber production and sustaining conservation values in natural tropical production forests. A systematic review protocol. Environmental Evidence 4: 8.  DOI: 10.1186/s13750-015-0034-7. C

Rendón Thompson, O.R., Paavola, J., Healey, J.R., Jones, J.P.G, Baker, T.R. & Torres, J. (2013). Reducing emissions from deforestation and forest degradation (REDD+): transaction costs of six Peruvian projects. Ecology and Society 18(1), 17 ( C

Pullin, A.S., Bangpan, M., Dalrymple, S., Dickson, K., Haddaway, N.R., Healey, J.R., Hauari, H., Hockley, N., Jones, J.P.G., Knight, T., Vigurs, C. & Oliver, S. (2013). Human well-being impacts of terrestrial protected areas. Environmental Evidence 2, 19.

Bowler, D.E., Buyung-Ali, L.M., Healey, J.R., Jones, J.P.G., Knight T.M. & Pullin, A.S. (2012). Does community forest management provide global environmental benefit and improve local welfare? Frontiers in Ecology and the Environment 10, 29-36. C


Tropical forest ecology and restoration

Poorter, L…, Healey, J.R.….. (2015). Diversity enhances carbon storage in tropical forests. Global Ecology and Biogeography 24, 1314-1328.  DOI:10.1111/geb.12364. C

Derroire, G., Powers, J.S., Hulshof, C.M., Cárdenas Varela, L.E. & Healey, J.R. (2018). Contrasting patterns of variation among and within species in leaf functional traits during tropical dry forest succession in Costa Rica. Scientific Reports 8, Article number 285.

Derroire, G., Balvanera, P., Castellanos-Castro, C., Decocq, G., Kennard, D., Lebrija-Trejos, E., Leiva, J., Oden, P.C., Powers, J., Rico-Gray, V., Tigabu, M. & Healey, J.R. (2016). Resilience of tropical dry forests – a meta-analysis of changes in species diversity and composition during secondary succession. Oikos 125, 1386-1397.  DOI: 10.1111/oik.03229.

Derroire, G., Coe, R. & Healey, J.R. (2016). Isolated trees as nuclei of regeneration in tropical pastures: testing the importance of niche-based and landscape factors. Journal of Vegetation Science 27, 679-691.  DOI: 10.1111/jvs.12404.

Derroire, G., Tigabu, M., Odén, P.C. & Healey, J.R. (2016). The effects of established trees on woody regeneration during secondary succession in tropical dry forests. Biotropica 48, 290-300.  DOI:10.1111/btp.12287.

Tanner, E.V.J., Rodriguez-Sanchez, F., Healey, J.R., Holdaway, R.J. & Bellingham, P.J. (2014). Long-term hurricane damage effects on tropical forest tree growth and mortality. Ecology 95, 2974-2983 (

Chai, S.-L., Healey, J.R. & Tanner, E.V.J. (2012). Evaluation of forest recovery over time and space using permanent plots monitored over 30 years in a Jamaican montane rain forest. PLOS ONE 7(11): e48859.  DOI:10.1371/journal.pone.0048859.



Rahman, S.A., Sunderland, T., Roshetko, J.M. & Healey, J.R. (2017). Facilitating smallholder tree farming in fragmented tropical landscapes: Challenges and potentials for sustainable land management. Journal of Environmental Management 198, 110-121 (

Rahman, S.A., Bredahl Jacobsen, J., Healey, J.R., Roshetko, J.M. & Sunderland, T. (2017). Finding alternatives to swidden agriculture: Does agroforestry improve livelihood options and reduce pressure on existing forest? Agroforestry Systems 91, 185-199.  (

Rahman, S., Sunderland, T., Kshatriya, M., Roshetko, J.M., Pagella, T. & Healey, J.R. (2016). Towards productive landscapes: Trade-offs in tree-cover and income across a matrix of smallholder agricultural land-use systems. Land Use Policy 58, 152-164 (

Rahman, S.A., Sunderland, T., Roshetko, J.M., Basuki, I. & Healey, J.R. (2016). Tree culture of smallholder farmers practicing agroforestry in Gunung Salak Valley, West Java, Indonesia. Small-scale Forestry 58, 152-164 (

Noponen, M.R.A., Healey, J.R., Soto, G. & Haggar, J.P. (2013). Sink or source – the potential of coffee agroforestry systems to sequester atmospheric CO2 into soil organic carbon. Agriculture, Ecosystems & Environment 175, 60-68. C

Noponen, M.R.A., Haggar, J.P., Edwards-Jones, G., Healey, J.R. (2013). Intensification of coffee systems can increase the effectiveness of REDD mechanisms. Agricultural Systems 119, 1-9.  C

Noponen, M.R.A., Edwards-Jones, G., Haggar, J.P., Soto, G., Attarzadeh, N. & Healey, J.R. (2012). Greenhouse gas emissions in coffee grown with differing input levels under conventional and organic management. Agriculture, Ecosystems & Environment 151, 6-15.  C


Temperate forest, heathland and peatland ecology and conservation

Avtzis D.N., Stara K., Sgardeli V., Betsis A., Diamandis S., Healey J.R., Kapsalis E., Kati V., Korakis G., Marini Govigli V., Monokrousos N., Muggia L., Nitsiakos V., Papadatou E., Papaioannou H., Rohrer A., Τsiakiris R., van Houtan K.S., Vokou D., Wong J. & Halley, JM. (2018). Quantifying the conservation value of sacred natural sites. Biological Conservation 222, 95-103.

Ford, H., Healey, J.R., Markesteijn, L. & Smith, A.R. (2018). How does grazing management influence the functional diversity of oak woodland ecosystems? A plant trait approach. Agriculture, Ecosystems and Environment 258, 154-161.

Gray, J. & Healey, J.R. (2016). Overwintering sites of ladybirds, shieldbugs and allied species in Hertfordshire woodland: how important is aspect? Transactions of the Hertfordshire Natural History Society 48 (1), 89-98.

Haddaway, N.R., Burden, A., Evans, C.D., Healey, J.R., Jones, D.L., Dalrymple, S.E. & Pullin, A.S. (2014). Evaluating effects of land management on greenhouse gas fluxes and carbon balances in boreo-temperate lowland peatland systems. Environmental Evidence 2014, 3: 5. DOI: 10.1186/2047-2382-3-5; C

Smith, A.R., Lukac, M., Hood, R., Healey, J.R., Miglietta, F. & Godbold, D.L. (2013). Elevated CO2 enrichment induces a differential biomass response in a mixed species temperate forest plantation. New Phytologist 198, 156-168. C

DeLuca, T.H., Zewdie, S.A., Zackrisson, O., Healey, J.R. & Jones, D.L. (2013). Bracken fern (Pteridium aquilinum L. Kuhn) promotes an open nitrogen cycle in heathland soils. Plant and Soil 367, 521-534. DOI: 10.1007/s11104-012-1484-0.


Tree disease bioeconomic modelling

Macpherson, M.F., Kleczkowski, A., Healey, J.R., Quine, C. & Hanley, N. (2017). The effects of invasive pests and diseases on strategies for forest diversification. Ecological Modelling 350, 87-99.

Macpherson, M.F., Kleczkowski, A., Healey, J.R. & Hanley, N. (2018). The effects of disease on optimal forest rotation: a generalisable analytical framework. Environmental and Resource Economics 70, 565-588.

Macpherson, M.F., Kleczkowski, A., Healey, J.R. & Hanley, N. (2017). Payment for multiple forest benefits alters the effect of tree disease on optimal forest rotation length. Ecological Economics 134, 82-94.

Sheremet, O., Healey, J.R., Quine, C. & Hanley, N. (2017). Public preferences and willingness to pay for forest disease control in the UK. Journal of Agricultural Economics 68, 781-800.


Nutrient cycling/resource use/sustainable production

Withers, P.J.A., Sylvester-Bradley, R., Jones, D.L., Healey, J.R. & Talboys, P.J. (2014). Feed the crop not the soil: rethinking phosphorus management in the food chain. Environmental Science & Technology 48, 6523-6530.

Talboys, P.J., Heppell, J., Roose, T., Healey, J.R., Jones, D.L. & Withers, P.J.A. (2016). Struvite: a slow-release fertiliser for sustainable phosphorus management? Plant and Soil 401, 109-123.  DOI:10.1007/s11104-015-2747-3.

Talboys, P.J., Healey, J.R., Withers, P.J.A. & Jones, D.L. (2014). Phosphate depletion modulates auxin transport in Triticum aestivium leading to altered root branching. Journal of Experimental Botany 65, 5023-5032.  DOI:10.1093/jxb/eru284.

Talboys, P.J., Owen, D.W., Healey, J.R., Withers, P.J.A. & Jones, D.L. (2014). Auxin secretion by Bacillus amyloliquefaciens FZB42 both stimulates root exudation and limits phosphorus uptake in Triticum aestivium. BMC Plant Biology 14: 51.  DOI:10.1186/1471-2229-14-51 (

Styles, D., Gibbons, J., Williams, A.P., Stichnothe, H., Chadwick, D.R. & Healey, J.R. (2015). Cattle feed or bioenergy? Consequential life cycle assessment of biogas feedstock options on dairy farms. Global Change Biology Bioenergy 7, 1034-1049.  DOI: 10.1111/gcbb.12189. C

Gibbons, J.M., Williamson, J.C., Williams, A.P., Withers, P.J.A., Hockley, N., Harris, I.M., Hughes, J.W., Taylor, R.L., Jones, D.L. & Healey, J.R. (2014). Sustainable nutrient management at field, farm and regional level: soil testing, nutrient budgets and the trade-off between lime application and greenhouse gas emissions. Agriculture, Ecosystems & Environment 188, 48-56. C

Ford, H., Smith, A., Pagella, T. & Healey, J. (2016). Trees, water storage and flooding in upland agricultural landscapes: why do we need to know more? Forestry & Timber News April 2016, 27-28.


Science-policy interface

O’Leary, B.C., Kvist, K., Bayliss, H.R., Derroire, G., Healey, J.R., Hughes, K., Kleinschroth, F., Sciberras, M., Woodcock, P. & Pullin, A.S. (2016). The reliability of evidence reviews in environmental science and conservation. Environmental Science and Policy 64, 75-82 (




Relena Ribbons. Seeing the forest for the trees: Tree species effects on soil microbial communities and nutrient cycling dynamics. Co-supervised with Morag McDonald and Andy Smith, SENRGY; Lars Vesterdal, Copenhagen University; and Cindy Prescott, University of British Columbia

Syed Rahman. Incorporation of trees in smallholder land use system: Farm characteristics, rates of return and policy issues influencing farmer adoption. Co-supervised with Jette Bredahl Jacobsen and Jens Friis Lund, University of Copenhagen; and Terry Sunderland, CIFOR

Lucia Morales Barquero. Beyond carbon accounting: A landscape perspective on measuring and monitoring tropical forest degradation. Co-supervised with Christoph Kleinn, University of Göttingen; Margaret Skutsch, National Autonomous University of Mexico

Fausto Bolom-Ton. Factors affecting variation in forest community characteristics and leaf-litter decomposition in tropical montane forest of Chiapas, Mexico: a functional ecology approach. Co-supervised with Bryan Finnegan, CATIE

Géraldine Derroire. Secondary succession in tropical dry forests: Drivers and mechanisms of forest regeneration. Co-supervised with Per Christer Oden and Mulualem Tigabu, Swedish University of Agricultural Sciences

Fritz Kleinschroth. Roads in the rainforests: Legacy of selective logging in Central Africa. Evaluating the temporal and spatial dynamics of logging road networks. Co-supervised with Sylvie Gourlet-Fleury and Plinio Sist, CIRAD, Montpellier.

Desalegn Desissa. Fragmentated forests in south-western Ethiopia: implications of land-use change on plant species composition and priorities for future conservation.

Jimena Esquivel Sheik. Plant traits and litter decomposition of tree species naturally regenerating in Central American pasturelands. Co-supervised with Bryan Finegan, CATIE

Martin Noponen. Carbon and economic performance of coffee agroforestry systems in Costa Rica and Nicaragua. Co-supervised with Gareth Edwards-Jones; and Jeremy Haggar and Gabriela Soto, CATIE

Deirdre Ninaber. Improved control of Rhododendron ponticum for environmental management. Co-supervised with Robert Brook

Sue Hearn. Perspectives in vegetation monitoring: an evaluation of approaches currently used in the UK. Co-supervised with Morag McDonald

Myers Paul. Composting of woodchip- and straw-based livestock bedding. Co-supervised with Davey Jones


Qualifications and Membership of Professional Bodies

BA Botany (University of Oxford)
PhD (University of Cambridge)
Member of the Institute of Chartered Foresters


Contact Info

Thoday Building, room S7


Tel: 01248 383703 (from U.K.)
+44 1248 383703 (International)


Google Scholar



Research and teaching focus: Forest ecology


For more information about my work click on "Research Interests" below

Teaching and Supervision

I am the organiser and lecturer for:

the final year undergraduate module DXX-3301 Forest Ecology    and

the MSc module DXX-4517 Forest Ecology and Resources.


I also contribute to:

the first year undergraduate modules DXX-1006 Making Snowdonia    and

DXX-1003 Forestry in the 21st Century ;

the second year undergraduate modules  DXX-2004 Silviculture and Inventory    and

DXX-2017 Forest Health ;

the MSc modules DDL-4202 Silviculture   and

DXX-4505 Natural Resources Management.


I also supervise several PhD students, whose details are given in the section "Research Interests".


Research areas and keywords


  • SD Forestry
  • QK Botany
  • GE Environmental Sciences

Research outputs (204)

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Prof. activities and awards (1)

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